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maravillas milkwort

desert milkwort, thorny milkwort

Habit Subshrubs, multi-stemmed, broomlike, 1.5–4 dm. Shrubs or subshrubs, single- to multi-stemmed, (1.5–)2–10(–12) dm.
Stems

usually erect, usually stiff, sometimes lax or sprawling, usually glaucous, especially proximally, glabrous.

sprawling to erect, densely pubescent to glabrate, hairs spreading and short.

Leaves

early deciduous; usually sessile, rarely subsessile;

blade scalelike, linear-subulate, lanceolate, or elliptic, 2(–3) × 0.5–1 mm, base and apex narrowly acute, surfaces pubescent, hairs incurved.

sessile or subsessile;

blade oblanceolate, narrowly obovate, or narrowly elliptic, 3–25 × 1–5 mm, base long-cuneate, apex rounded or acute, surfaces usually densely pubescent, rarely subglabrous, hairs spreading and short.

Racemes

terminal, often also appearing axillary (from branches proximal to racemes of major branches with vegetative portions highly reduced), 2–10(–15) × 0.8–1.9 cm;

rachis not thorn-tipped;

peduncle to 2 cm, sometimes vestigial, especially on reduced axillary racemes;

bracts mostly deciduous, rarely persistent, lanceolate, narrowly ovate, or linear.

terminal, sometimes appearing axillary if poorly developed, sometimes aggregated into pseudopanicles or reduced and appearing fasciculate, 0.5–2.5 ×0.6–2 cm;

rachis thorn-tipped;

peduncle 0–0.2(–0.5) cm, often poorly developed;

bracts deciduous, lanceolate.

Pedicels

1.5–3.2(–3.6) mm, glabrous.

1.5–4(–5.8) mm, usually shorter than flowers, pubescent, sometimes sparsely so.

Flowers

pink, keel green to yellow distally, (3–)3.4–5 mm;

upper sepal persistent, others deciduous, ovate to elliptic, lower sepals ovate or elliptic to narrowly obovate, 1.5–2.8 mm, glabrous, margins sparsely ciliate proximally;

wings obovate, (2.7–)3.5–4.7 × (1.5–)1.8–2.8 mm, glabrous, margins sometimes sparsely ciliate proximally;

keel (2.5–)2.7–3.5 mm, sac incurved-puberulent in distal 1/2, beak bluntly rounded, 0.3–0.8 × 0.3–0.6 mm, pubescent.

cream to yellowish green, wings cream to greenish, upper petals often purple-tipped, distal keel often dark yellow to green, (3–)3.5–5(–5.3) mm;

sepals deciduous, ovate to elliptic, 1.6–3.5 mm, spreading-pubescent, margins usually ciliate;

wings obovate, 3–5 × 2–3 mm, glabrous or sparsely pubescent;

keel 2.7–3.8 mm, sac glabrous, beak absent or obscure and bluntly rounded, to 0.7 × 0.5 mm, glabrous;

stamens rarely 9.

Capsules

obovoid, usually narrowly so, (2.6–)3.3–4.4 × 1.8–2.6 mm, base cuneate, margins with very narrow and even wing, glabrous or sparsely pubescent apically.

ellipsoid or slightly obovoid, (3–)4–6 × 2.5–4 mm, base rounded or, sometimes, cuneate, margins with very narrow and even wing, glabrous.

Seeds

2.3–2.9 mm, pubescent, usually more sparsely pubescent to often subglabrous in distal 1/5–1/2 (sometimes evenly pubescent throughout);

aril 0.6–1.1 mm, lobes to 1/3 length of seed.

3.2–4.2 mm, pubescent;

aril 1–1.7 mm, lobes to 1/3 length of seed.

2n

= 18 (36).

= 18.

Rhinotropis maravillasensis

Rhinotropis acanthoclada

Phenology Flowering spring–fall. Flowering (early spring–)spring–summer(–late fall).
Habitat Crevices of limestone rocks and cliffs in desert and semidesert canyons and hills. Usually on loose silts or sands derived from limestone, granite, sandstone, or gypsum in open places or slopes in desert scrub or juniper woodlands.
Elevation 400–900 m. (1300–3000 ft.) 800–1800 m. (2600–5900 ft.)
Distribution
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

In the flora area, Rhinotropis maravillasensis occurs along the Rio Grande in Brewster and Terrell counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In northern Arizona and southern Utah, Rhinotropis acanthoclada overlaps geographically with R. intermontana and tetraploid hybrids are known (T. L. Wendt 1978, 1979).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
Synonyms Polygala maravillasensis Polygalaacanthoclada a.
Name authority (Correll) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011)
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