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shrubby milkwort

Habit Herbs, multi-stemmed, 0.3–3(–3.5) dm (rarely straggling to 10 dm). Herbs or subshrubs, multi-stemmed, often loosely mat-forming to 2.5 dm diam., 0.1–0.5 dm.
Stems

decumbent to erect, usually pubescent, rarely glabrous, hairs spreading or incurved.

prostrate to laxly erect, glabrous or very sparsely pubescent, hairs incurved.

Leaves

subsessile to petiolate, petiole to 1(–1.5) mm;

blade elliptic to linear, lanceolate, ovate, obovate, or scalelike, (3–)4–41 × (0.5–)1–12(–18) mm, base rounded to cuneate, apex obtuse to rounded, surfaces pubescent or glabrous, hairs incurved or spreading.

sessile or subsessile;

blade elliptic, obovate, ovate, or scalelike, (1.5–)2–5.9(–8) × 0.8–3.5 mm, base cuneate or rounded, apex acute, surfaces sparsely pubescent to subglabrous, hairs incurved.

Racemes

terminal, usually leaf-opposed, often also from near base of plant, these usually with chasmogamous flowers, occasionally bearing reduced, beakless cleistogamous or semi-cleistogamous flowers, rarely with cleistogamous or semi-cleistogamous flowers throughout, 1–12(–15) × 0.3–1.5 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts usually persistent, ovate, lanceolate, or elliptic.

terminal, sometimes appearing axillary (from branches proximal to racemes of major branches with vegetative portions highly reduced), reduced to 1–3(–5) flowers, 0.5–1 × 0.7–1.9 cm;

peduncle 0–0.2 cm;

rachis not thorn-tipped;

bracts persistent, ovate, lanceolate, or linear.

Pedicels

1–4.5 mm, pubescent.

(0.6–)1–3(–3.6) mm, usually glabrous, rarely sparsely pubescent.

Flowers

usually pink to purple, rarely white, keel yellowish distally, wings pink or rose, (3.7–)4–7.4(–7.7) mm;

upper sepal persistent, other sepals deciduous, upper sepal ovate, 1.7–4.5(–5.2) mm, lower sepals lanceolate to obovate, (1.3–)1.6–3.5(–3.8) mm, pubescent or glabrous;

wings obovate to oblong-obovate, 3–6.4(–7.2) × (1.2–)1.4–3.2 mm, glabrous or pubescent;

keel (2.7–)3.1–6.2 mm, sac glabrous or with scattered hairs, beak linear (or bluntly rounded), (0–)0.5–2 × (0–)0.2–0.6 mm, glabrous or pubescent.

pink or purple with pale margins, keel sometimes cream, distally greenish yellow, (2.4–)2.9–5.1(–5.4) mm;

sepals: lateral ones deciduous, elliptic, ovate, or obovate, (1.2–)1.6–3(–3.2) mm, upper sepal persistent, ovate, (1–)1.2–2.4 mm, glabrous, margins sometimes sparsely ciliate proximally;

wings obovate, (2.2–)2.5–4.6(–5.1) × 1.8–2.8 mm, glabrous or sparsely pubescent proximally;

keel (1.7–)2.3–3.6 mm, sac incurved-puberulent in distal 1/2, beak obscure, deltate, bluntly rounded, or linear to oblong, (0–)0.1–0.7 × (0–)0.1–0.5 mm, glabrous.

Capsules

ellipsoid, oblong, slightly ovoid, or obovoid, 3.3–6(–6.8) × 2–4 mm, base rounded to subtruncate, often oblique, margins with narrow wing or not winged, usually pubescent, rarely subglabrous.

ellipsoid-obovoid, sometimes broadly so, 1.9–3.6 × 1.6–2.9 mm, base cuneate to rounded, margins narrowly and evenly winged or slightly expanded apically, glabrous or sparsely pubescent apically.

Seeds

2.8–4.3 mm, pubescent;

aril 0.7–2.5 mm, lobes to 3/4 length of seed.

(1.7–)1.9–3.2 mm, body, excluding aril and pubescence, 1–1.9 mm, densely pubescent;

aril (0.3–)0.4–0.7 mm, lobes often highly reduced, nearly absent to 1/6 length of seed.

Rhinotropis lindheimeri

Rhinotropis rimulicola

Distribution
sw United States; sc United States; n Mexico
[BONAP county map]
sw United States; sc United States
[BONAP county map]
Discussion

Varieties 3 (2 in the flora).

Variety eucosma (S. F. Blake) T. Wendt is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually with spreading hairs, hairs rarely somewhat crisped, mostly 0.3–0.5 mm; leaf blades usually elliptic, ovate, or obovate proximally, distally becoming narrowly so, venation usually prominently reticulate, surfaces pubescent (not glabrous); keel sacs glabrous or with scattered, spreading hairs proximally, hairs not incurved in distal 1/2.
var. lindheimeri
1. Stems usually with incurved hairs, hairs rarely irregularly spreading, 0.07–0.15 mm, rarely glabrous; leaf blades lanceolate, linear, or scalelike to elliptic, ovate, or obovate, venation usually not prominently reticulate (usually midvein prominent abaxially, occasionally reticulate), surfaces pubescent or glabrous; keel sacs glabrous or, rarely, with incurved hairs in distal 1/2.
var. parvifolia
1. Keel petals with deltate or rounded beaks (often obscure), (0–)0.1–0.3(–0.5) mm, base width equal to or greater than length; seeds (1.7–)1.9–2.7 mm; sepals eciliate or, infrequently, with a few proximal cilia.
var. rimulicola
1. Keel petals with linear to oblong beaks, 0.3–0.7 mm, base width usually less than 2/3 length; seeds 2.8–3.2 mm; sepals sparsely ciliate in proximal 1/2.
var. mescalerorum
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. maravillasensis, R. nitida, R. nudata, R. rusbyi, R. subspinosa
Subordinate taxa
R. lindheimeri var. lindheimeri, R. lindheimeri var. parvifolia
R. rimulicola var. mescalerorum, R. rimulicola var. rimulicola
Synonyms Polygalalindheimeri a. Polygalarimulicola steyermark
Name authority (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (Steyermark) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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