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shrubby milkwort

small-flower milkwort

Habit Herbs, multi-stemmed, 0.3–3(–3.5) dm (rarely straggling to 10 dm). Subshrubs, multi-stemmed, broomlike, 1.5–4(–5) dm.
Stems

decumbent to erect, usually pubescent, rarely glabrous, hairs spreading or incurved.

usually erect to decumbent or procumbent, usually sparsely pubescent, rarely glabrate, hairs incurved.

Leaves

subsessile to petiolate, petiole to 1(–1.5) mm;

blade elliptic to linear, lanceolate, ovate, obovate, or scalelike, (3–)4–41 × (0.5–)1–12(–18) mm, base rounded to cuneate, apex obtuse to rounded, surfaces pubescent or glabrous, hairs incurved or spreading.

larger ones usually early deciduous;

sessile or petiolate, petiole to 1 mm;

blade linear, lanceolate, narrowly elliptic, obovate, or scalelike, 1–4(–15) × 0.5–1(–3.7) mm, base cuneate, apex acute, surfaces glabrous or sparsely pubescent, hairs incurved.

Racemes

terminal, usually leaf-opposed, often also from near base of plant, these usually with chasmogamous flowers, occasionally bearing reduced, beakless cleistogamous or semi-cleistogamous flowers, rarely with cleistogamous or semi-cleistogamous flowers throughout, 1–12(–15) × 0.3–1.5 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts usually persistent, ovate, lanceolate, or elliptic.

terminal, often also appearing axillary (from branches proximal to racemes of major branches with vegetative portions highly reduced), 2–10(–15) ×0.8–1.5 cm;

rachis not thorn-tipped;

peduncle 0–2 cm;

bracts persistent, ovate to lanceolate.

Pedicels

1–4.5 mm, pubescent.

(1.3–)1.6–3.6 mm, usually pubescent, rarely subglabrous.

Flowers

usually pink to purple, rarely white, keel yellowish distally, wings pink or rose, (3.7–)4–7.4(–7.7) mm;

upper sepal persistent, other sepals deciduous, upper sepal ovate, 1.7–4.5(–5.2) mm, lower sepals lanceolate to obovate, (1.3–)1.6–3.5(–3.8) mm, pubescent or glabrous;

wings obovate to oblong-obovate, 3–6.4(–7.2) × (1.2–)1.4–3.2 mm, glabrous or pubescent;

keel (2.7–)3.1–6.2 mm, sac glabrous or with scattered hairs, beak linear (or bluntly rounded), (0–)0.5–2 × (0–)0.2–0.6 mm, glabrous or pubescent.

usually white or cream, often with purplish center stripe, rarely pink, 3–5 mm;

upper sepal persistent, others deciduous, ovate to elliptic or obovate, 1.4–3(–3.4) mm, pubescent;

wings obovate to elliptic, (2.5–)3–4.6 × (1.2–)1.5–3 mm, usually glabrous, rarely sparsely pubescent;

keel (2.2–)2.5–4 mm, sac incurved-puberulent in distal 1/2, beak bluntly rounded to oblong, (0.3–)0.4–1.1 × 0.2–0.5(–0.7) mm, pubescent.

Capsules

ellipsoid, oblong, slightly ovoid, or obovoid, 3.3–6(–6.8) × 2–4 mm, base rounded to subtruncate, often oblique, margins with narrow wing or not winged, usually pubescent, rarely subglabrous.

ovoid to broadly ellipsoid, 2.5–4(–4.5) × (2.1–)2.3–3.2(–3.4) mm, base rounded or subtruncate, margins with very narrow and even wing, usually pubescent, sometimes subglabrous proximally.

Seeds

2.8–4.3 mm, pubescent;

aril 0.7–2.5 mm, lobes to 3/4 length of seed.

2.3–3(–3.4) mm, usually evenly pubescent;

aril (0.8–)1–1.6 mm, lobes to 1/2 length of seed.

2n

= 18.

Rhinotropis lindheimeri

Rhinotropis nudata

Phenology Flowering spring–late fall.
Habitat Mostly on limestone, also on gypsum, sandstone, or tuff, rocky slopes in desert scrub, chaparral, or mixed woodlands (pinyon, juniper, or oak).
Elevation 1200–1700 m. (3900–5600 ft.)
Distribution
sw United States; sc United States; n Mexico
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas)
[BONAP county map]
Discussion

Varieties 3 (2 in the flora).

Variety eucosma (S. F. Blake) T. Wendt is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rhinotropis nudata occurs in Brewster and Presidio counties.

The name Polygala minutifolia Rose was misapplied to this taxon by D. S. Correll and M. C. Johnston (1970).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually with spreading hairs, hairs rarely somewhat crisped, mostly 0.3–0.5 mm; leaf blades usually elliptic, ovate, or obovate proximally, distally becoming narrowly so, venation usually prominently reticulate, surfaces pubescent (not glabrous); keel sacs glabrous or with scattered, spreading hairs proximally, hairs not incurved in distal 1/2.
var. lindheimeri
1. Stems usually with incurved hairs, hairs rarely irregularly spreading, 0.07–0.15 mm, rarely glabrous; leaf blades lanceolate, linear, or scalelike to elliptic, ovate, or obovate, venation usually not prominently reticulate (usually midvein prominent abaxially, occasionally reticulate), surfaces pubescent or glabrous; keel sacs glabrous or, rarely, with incurved hairs in distal 1/2.
var. parvifolia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. maravillasensis, R. nitida, R. rimulicola, R. rusbyi, R. subspinosa
Subordinate taxa
R. lindheimeri var. lindheimeri, R. lindheimeri var. parvifolia
Synonyms Polygalalindheimeri a. Polygalanudata brandegee
Name authority (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (Brandegee) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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