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shrubby milkwort

shining milkwort

Habit Herbs, multi-stemmed, 0.3–3(–3.5) dm (rarely straggling to 10 dm). Herbs, multi-stemmed, 0.5–2 dm; roots with bright red to orange cortex, loosely exfoliating in thin layers.
Stems

decumbent to erect, usually pubescent, rarely glabrous, hairs spreading or incurved.

spreading to erect, pubescent, hairs incurved and appressed.

Leaves

subsessile to petiolate, petiole to 1(–1.5) mm;

blade elliptic to linear, lanceolate, ovate, obovate, or scalelike, (3–)4–41 × (0.5–)1–12(–18) mm, base rounded to cuneate, apex obtuse to rounded, surfaces pubescent or glabrous, hairs incurved or spreading.

subsessile or petiolate, petiole to 3 mm;

blade ovate, lanceolate, elliptic, or linear, 5–35(–40) × 1–9 mm, base rounded to cuneate, apex acute to rounded or acuminate, surfaces subglabrous or pubescent, hairs incurved.

Racemes

terminal, usually leaf-opposed, often also from near base of plant, these usually with chasmogamous flowers, occasionally bearing reduced, beakless cleistogamous or semi-cleistogamous flowers, rarely with cleistogamous or semi-cleistogamous flowers throughout, 1–12(–15) × 0.3–1.5 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts usually persistent, ovate, lanceolate, or elliptic.

terminal, usually leaf-opposed, sometimes near stem base, 0.8–5(–6) × 1–2.5 cm;

rachis not thorn-tipped;

peduncle 0–0.5 cm;

bracts persistent, elliptic, ovate, or lanceolate.

Pedicels

1–4.5 mm, pubescent.

1.5–4 mm, pubescent or glabrous.

Flowers

usually pink to purple, rarely white, keel yellowish distally, wings pink or rose, (3.7–)4–7.4(–7.7) mm;

upper sepal persistent, other sepals deciduous, upper sepal ovate, 1.7–4.5(–5.2) mm, lower sepals lanceolate to obovate, (1.3–)1.6–3.5(–3.8) mm, pubescent or glabrous;

wings obovate to oblong-obovate, 3–6.4(–7.2) × (1.2–)1.4–3.2 mm, glabrous or pubescent;

keel (2.7–)3.1–6.2 mm, sac glabrous or with scattered hairs, beak linear (or bluntly rounded), (0–)0.5–2 × (0–)0.2–0.6 mm, glabrous or pubescent.

pink to cream, wings sometimes greenish, (4.4–)5–7.5 mm;

upper sepal persistent, others deciduous, ovate, lower sepals ovate to elliptic or obovate, (1.6–)1.9–4.1 mm, pubescent or glabrous;

wings obovate to elliptic, 3–7 × 1.4–3.2 mm, pubescent or glabrous;

keel 2.7–6.1 mm, sac usually glabrous, rarely sparsely incurved-pubescent distally, beak oblong or bluntly rounded, (0.2–)0.6–1.7 × 0.2–0.8 mm (rarely absent in var. tamaulipana), glabrous or pubescent.

Capsules

ellipsoid, oblong, slightly ovoid, or obovoid, 3.3–6(–6.8) × 2–4 mm, base rounded to subtruncate, often oblique, margins with narrow wing or not winged, usually pubescent, rarely subglabrous.

ellipsoid, sometimes broadly so, 3.7–6.2 × 2.4–4.3 mm, base subtruncate to acute, margins with very narrow wing or not winged, pubescent.

Seeds

2.8–4.3 mm, pubescent;

aril 0.7–2.5 mm, lobes to 3/4 length of seed.

2.7–4.9 mm, densely pubescent;

aril 1.3–2.6 mm, lobes to 1/2 length of seed.

Rhinotropis lindheimeri

Rhinotropis nitida

Distribution
sw United States; sc United States; n Mexico
[BONAP county map]
n Mexico; Texas
[BONAP county map]
Discussion

Varieties 3 (2 in the flora).

Variety eucosma (S. F. Blake) T. Wendt is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (2 in the flora).

Rhinotropis nitida is closely related to R. lindheimeri, with seven varieties recognized between the two. According to T. L. Wendt (1978), although the differences between the two species are fairly small, the recognition of a single species would obscure the differences in the evolution of several superficially similar taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually with spreading hairs, hairs rarely somewhat crisped, mostly 0.3–0.5 mm; leaf blades usually elliptic, ovate, or obovate proximally, distally becoming narrowly so, venation usually prominently reticulate, surfaces pubescent (not glabrous); keel sacs glabrous or with scattered, spreading hairs proximally, hairs not incurved in distal 1/2.
var. lindheimeri
1. Stems usually with incurved hairs, hairs rarely irregularly spreading, 0.07–0.15 mm, rarely glabrous; leaf blades lanceolate, linear, or scalelike to elliptic, ovate, or obovate, venation usually not prominently reticulate (usually midvein prominent abaxially, occasionally reticulate), surfaces pubescent or glabrous; keel sacs glabrous or, rarely, with incurved hairs in distal 1/2.
var. parvifolia
1. Stems spreading to erect, hairs mostly 0.15–0.3 mm, usually spreading to loosely incurved, tips not close to stem surface; leaf blades elliptic to ovate in proximal 1/3 of stem, length 1.5–3 times width, distal leaves similar or somewhat narrower, not lanceolate or linear; pedicels (2–)2.5–4 mm.
var. goliadensis
1. Stems erect, hairs mostly 0.07–0.1(–0.15) mm, closely incurved-appressed, very close to stem, tips touching stem surface; leaf blades narrowly elliptic to lanceolate or linear, including basal ones, length at least 5 times width, or when leaves in proximal 1/3 of stem are broader, then distal leaves lanceolate-elliptic, much narrower than proximal ones; pedicels 1.5–3 mm.
var. tamaulipana
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. maravillasensis, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
Subordinate taxa
R. lindheimeri var. lindheimeri, R. lindheimeri var. parvifolia
R. nitida var. goliadensis, R. nitida var. tamaulipana
Synonyms Polygalalindheimeri a. Polygalanitida brandegee
Name authority (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (Brandegee) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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