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shrubby milkwort

maravillas milkwort

Habit Herbs, multi-stemmed, 0.3–3(–3.5) dm (rarely straggling to 10 dm). Subshrubs, multi-stemmed, broomlike, 1.5–4 dm.
Stems

decumbent to erect, usually pubescent, rarely glabrous, hairs spreading or incurved.

usually erect, usually stiff, sometimes lax or sprawling, usually glaucous, especially proximally, glabrous.

Leaves

subsessile to petiolate, petiole to 1(–1.5) mm;

blade elliptic to linear, lanceolate, ovate, obovate, or scalelike, (3–)4–41 × (0.5–)1–12(–18) mm, base rounded to cuneate, apex obtuse to rounded, surfaces pubescent or glabrous, hairs incurved or spreading.

early deciduous; usually sessile, rarely subsessile;

blade scalelike, linear-subulate, lanceolate, or elliptic, 2(–3) × 0.5–1 mm, base and apex narrowly acute, surfaces pubescent, hairs incurved.

Racemes

terminal, usually leaf-opposed, often also from near base of plant, these usually with chasmogamous flowers, occasionally bearing reduced, beakless cleistogamous or semi-cleistogamous flowers, rarely with cleistogamous or semi-cleistogamous flowers throughout, 1–12(–15) × 0.3–1.5 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts usually persistent, ovate, lanceolate, or elliptic.

terminal, often also appearing axillary (from branches proximal to racemes of major branches with vegetative portions highly reduced), 2–10(–15) × 0.8–1.9 cm;

rachis not thorn-tipped;

peduncle to 2 cm, sometimes vestigial, especially on reduced axillary racemes;

bracts mostly deciduous, rarely persistent, lanceolate, narrowly ovate, or linear.

Pedicels

1–4.5 mm, pubescent.

1.5–3.2(–3.6) mm, glabrous.

Flowers

usually pink to purple, rarely white, keel yellowish distally, wings pink or rose, (3.7–)4–7.4(–7.7) mm;

upper sepal persistent, other sepals deciduous, upper sepal ovate, 1.7–4.5(–5.2) mm, lower sepals lanceolate to obovate, (1.3–)1.6–3.5(–3.8) mm, pubescent or glabrous;

wings obovate to oblong-obovate, 3–6.4(–7.2) × (1.2–)1.4–3.2 mm, glabrous or pubescent;

keel (2.7–)3.1–6.2 mm, sac glabrous or with scattered hairs, beak linear (or bluntly rounded), (0–)0.5–2 × (0–)0.2–0.6 mm, glabrous or pubescent.

pink, keel green to yellow distally, (3–)3.4–5 mm;

upper sepal persistent, others deciduous, ovate to elliptic, lower sepals ovate or elliptic to narrowly obovate, 1.5–2.8 mm, glabrous, margins sparsely ciliate proximally;

wings obovate, (2.7–)3.5–4.7 × (1.5–)1.8–2.8 mm, glabrous, margins sometimes sparsely ciliate proximally;

keel (2.5–)2.7–3.5 mm, sac incurved-puberulent in distal 1/2, beak bluntly rounded, 0.3–0.8 × 0.3–0.6 mm, pubescent.

Capsules

ellipsoid, oblong, slightly ovoid, or obovoid, 3.3–6(–6.8) × 2–4 mm, base rounded to subtruncate, often oblique, margins with narrow wing or not winged, usually pubescent, rarely subglabrous.

obovoid, usually narrowly so, (2.6–)3.3–4.4 × 1.8–2.6 mm, base cuneate, margins with very narrow and even wing, glabrous or sparsely pubescent apically.

Seeds

2.8–4.3 mm, pubescent;

aril 0.7–2.5 mm, lobes to 3/4 length of seed.

2.3–2.9 mm, pubescent, usually more sparsely pubescent to often subglabrous in distal 1/5–1/2 (sometimes evenly pubescent throughout);

aril 0.6–1.1 mm, lobes to 1/3 length of seed.

2n

= 18 (36).

Rhinotropis lindheimeri

Rhinotropis maravillasensis

Phenology Flowering spring–fall.
Habitat Crevices of limestone rocks and cliffs in desert and semidesert canyons and hills.
Elevation 400–900 m. (1300–3000 ft.)
Distribution
sw United States; sc United States; n Mexico
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Varieties 3 (2 in the flora).

Variety eucosma (S. F. Blake) T. Wendt is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the flora area, Rhinotropis maravillasensis occurs along the Rio Grande in Brewster and Terrell counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually with spreading hairs, hairs rarely somewhat crisped, mostly 0.3–0.5 mm; leaf blades usually elliptic, ovate, or obovate proximally, distally becoming narrowly so, venation usually prominently reticulate, surfaces pubescent (not glabrous); keel sacs glabrous or with scattered, spreading hairs proximally, hairs not incurved in distal 1/2.
var. lindheimeri
1. Stems usually with incurved hairs, hairs rarely irregularly spreading, 0.07–0.15 mm, rarely glabrous; leaf blades lanceolate, linear, or scalelike to elliptic, ovate, or obovate, venation usually not prominently reticulate (usually midvein prominent abaxially, occasionally reticulate), surfaces pubescent or glabrous; keel sacs glabrous or, rarely, with incurved hairs in distal 1/2.
var. parvifolia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
Subordinate taxa
R. lindheimeri var. lindheimeri, R. lindheimeri var. parvifolia
Synonyms Polygalalindheimeri a. Polygala maravillasensis
Name authority (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (Correll) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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