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shrubby milkwort

intermountain milkwort

Habit Herbs, multi-stemmed, 0.3–3(–3.5) dm (rarely straggling to 10 dm). Subshrubs or shrubs, multi-stemmed, sometimes mat-forming, 1.5–10 dm.
Stems

decumbent to erect, usually pubescent, rarely glabrous, hairs spreading or incurved.

erect to sprawling, densely pubescent or glabrate, with dense, matted or shaggy tomentum, hairs appressed, incurved, or, occasionally, irregularly spreading.

Leaves

subsessile to petiolate, petiole to 1(–1.5) mm;

blade elliptic to linear, lanceolate, ovate, obovate, or scalelike, (3–)4–41 × (0.5–)1–12(–18) mm, base rounded to cuneate, apex obtuse to rounded, surfaces pubescent or glabrous, hairs incurved or spreading.

sessile or subsessile, rarely with narrow, petiolelike base to 1(–2) mm;

blade linear to oblanceolate or obovate, (3–)4–20(–25) × 0.8–3(–3.5) mm, base long-cuneate, apex rounded to acute, surfaces densely pubescent, hairs incurved.

Racemes

terminal, usually leaf-opposed, often also from near base of plant, these usually with chasmogamous flowers, occasionally bearing reduced, beakless cleistogamous or semi-cleistogamous flowers, rarely with cleistogamous or semi-cleistogamous flowers throughout, 1–12(–15) × 0.3–1.5 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts usually persistent, ovate, lanceolate, or elliptic.

terminal, sometimes aggregated into pseudopanicles or reduced and appearing fasciculate, 1.5 × 0.7–1.3 cm;

rachis thorn-tipped;

peduncle 0–0.1 cm;

bracts deciduous, lanceolate or ovate.

Pedicels

1–4.5 mm, pubescent.

(2.5–)3–7(–9) mm, glabrous.

Flowers

usually pink to purple, rarely white, keel yellowish distally, wings pink or rose, (3.7–)4–7.4(–7.7) mm;

upper sepal persistent, other sepals deciduous, upper sepal ovate, 1.7–4.5(–5.2) mm, lower sepals lanceolate to obovate, (1.3–)1.6–3.5(–3.8) mm, pubescent or glabrous;

wings obovate to oblong-obovate, 3–6.4(–7.2) × (1.2–)1.4–3.2 mm, glabrous or pubescent;

keel (2.7–)3.1–6.2 mm, sac glabrous or with scattered hairs, beak linear (or bluntly rounded), (0–)0.5–2 × (0–)0.2–0.6 mm, glabrous or pubescent.

cream or greenish, (2.5–)3–4.7(–5.2) mm;

sepals deciduous, ovate or elliptic, 1.3–3.3 mm, glabrous or with few incurved hairs subapically, margins sparsely ciliate;

wings obovate, 2.5–4.9 × 1.5–3 mm, glabrous or sparsely pubescent subapically;

keel (2–)2.5–3.4 mm, sac glabrous or appressed-pubescent in upper part, beak mostly absent, when present, a bluntly rounded projection, 0(–0.5) × 0(–0.5) mm, glabrous or pubescent.

Capsules

ellipsoid, oblong, slightly ovoid, or obovoid, 3.3–6(–6.8) × 2–4 mm, base rounded to subtruncate, often oblique, margins with narrow wing or not winged, usually pubescent, rarely subglabrous.

broadly ellipsoid, ovoid, or subglobose, 3.5–5.8 × 3.3–4.6 mm, base truncate to rounded, margins with narrow and even wing, glabrous.

Seeds

2.8–4.3 mm, pubescent;

aril 0.7–2.5 mm, lobes to 3/4 length of seed.

2.8–4.2 mm, sparsely pubescent to subglabrous;

aril 1.2–2.3 mm, lobes to 1/3 length of seed.

2n

= 18.

Rhinotropis lindheimeri

Rhinotropis intermontana

Phenology Flowering spring–early summer(–fall).
Habitat Sandy, gravelly, or loose silt flats, slopes, dunes, ridges, and badlands of diverse parent materials in open desert scrub or mountain slopes in pinyon-juniper-sagebrush woodlands, sagebrush scrub.
Elevation 600–3000 m. (2000–9800 ft.)
Distribution
sw United States; sc United States; n Mexico
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

Varieties 3 (2 in the flora).

Variety eucosma (S. F. Blake) T. Wendt is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rhinotropis intermontana is named for its distribution in the Intermountain region of the United States, which is bounded by the Rocky Mountains on the east, the Sierra Nevada and Cascade Range on the west, and the Mojave Desert to the south.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually with spreading hairs, hairs rarely somewhat crisped, mostly 0.3–0.5 mm; leaf blades usually elliptic, ovate, or obovate proximally, distally becoming narrowly so, venation usually prominently reticulate, surfaces pubescent (not glabrous); keel sacs glabrous or with scattered, spreading hairs proximally, hairs not incurved in distal 1/2.
var. lindheimeri
1. Stems usually with incurved hairs, hairs rarely irregularly spreading, 0.07–0.15 mm, rarely glabrous; leaf blades lanceolate, linear, or scalelike to elliptic, ovate, or obovate, venation usually not prominently reticulate (usually midvein prominent abaxially, occasionally reticulate), surfaces pubescent or glabrous; keel sacs glabrous or, rarely, with incurved hairs in distal 1/2.
var. parvifolia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. intermontana, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. lindheimeri, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
Subordinate taxa
R. lindheimeri var. lindheimeri, R. lindheimeri var. parvifolia
Synonyms Polygalalindheimeri a. Polygalaintermontana t.
Name authority (A. Gray) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011) (T. Wendt) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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