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California milkwort

intermountain milkwort

Habit Herbs, sometimes suffrutescent, multi-stemmed, often forming a ground cover, 0.5–3.5 dm. Subshrubs or shrubs, multi-stemmed, sometimes mat-forming, 1.5–10 dm.
Stems

laxly erect, decumbent, or prostrate, pubescent to subglabrous, hairs incurved.

erect to sprawling, densely pubescent or glabrate, with dense, matted or shaggy tomentum, hairs appressed, incurved, or, occasionally, irregularly spreading.

Leaves

sessile or subsessile;

blade ovate, elliptic, or obovate, 7–50(–60) × 3–20(–26) mm, base usually rounded to acute, sometimes cuneate, apex rounded to acute, surfaces pubescent, hairs incurved.

sessile or subsessile, rarely with narrow, petiolelike base to 1(–2) mm;

blade linear to oblanceolate or obovate, (3–)4–20(–25) × 0.8–3(–3.5) mm, base long-cuneate, apex rounded to acute, surfaces densely pubescent, hairs incurved.

Racemes

terminal or leaf-opposed, 1–4(–5) × 1.8–3 cm;

rachis not thorn-tipped;

peduncle 0–1 cm;

bracts early deciduous, linear to lanceolate.

terminal, sometimes aggregated into pseudopanicles or reduced and appearing fasciculate, 1.5 × 0.7–1.3 cm;

rachis thorn-tipped;

peduncle 0–0.1 cm;

bracts deciduous, lanceolate or ovate.

Pedicels

(2.5–)3.5–8.5 mm, sparsely pubescent or glabrous.

(2.5–)3–7(–9) mm, glabrous.

Flowers

usually pink, rarely white, keel distally yellow (fading white), (2.5–)9–14.5 mm, cleistogamous and semi-cleistogamous flowers mostly 2.5–5 mm, intergrading with chasmogamous flowers;

sepals deciduous, elliptic, 4–6.5 mm, pubescent or glabrous;

wings obovate, (7.5–)8–12 × 2.5–6 mm, glabrous or sparsely pubescent;

keel (7–)8–11 mm, sac glabrous (sometimes proximally ciliate), beak oblong, (1.2–)1.6–3 × 0.7–1 mm (mostly absent in cleistogamous flowers), usually notched or contorted abaxially, rarely subentire, pubescent.

cream or greenish, (2.5–)3–4.7(–5.2) mm;

sepals deciduous, ovate or elliptic, 1.3–3.3 mm, glabrous or with few incurved hairs subapically, margins sparsely ciliate;

wings obovate, 2.5–4.9 × 1.5–3 mm, glabrous or sparsely pubescent subapically;

keel (2–)2.5–3.4 mm, sac glabrous or appressed-pubescent in upper part, beak mostly absent, when present, a bluntly rounded projection, 0(–0.5) × 0(–0.5) mm, glabrous or pubescent.

Capsules

ellipsoid to ovoid, 7.3–10.5 × 4.5–7 mm, in cleistogamous and semi-cleistogamous flowers 4.5–8 mm, base obtuse, rounded, or subtruncate, margins with narrow, entire or slightly erose wing, glabrous, margins sometimes ciliolate.

broadly ellipsoid, ovoid, or subglobose, 3.5–5.8 × 3.3–4.6 mm, base truncate to rounded, margins with narrow and even wing, glabrous.

Seeds

3.5–6 mm, densely pubescent;

aril 1.7–4 mm, less than 1/2 length of seed.

2.8–4.2 mm, sparsely pubescent to subglabrous;

aril 1.2–2.3 mm, lobes to 1/3 length of seed.

Cleistogamous

or semi-cleistogamous flowers often present terminally, on much reduced scale-leaved lateral branches from proximal (or distal) leaf axils, or terminally on leafy branches that are often leaf-opposed.

2n

= 18.

= 18.

Rhinotropis californica

Rhinotropis intermontana

Phenology Flowering spring–summer. Flowering spring–early summer(–fall).
Habitat Rocky or clay soils, deep duff, rich soils, serpentine soils, slopes or drainages, full sun to deep shade, open habitat, chaparral, mixed evergreen forests, oak woodlands, coniferous forests. Sandy, gravelly, or loose silt flats, slopes, dunes, ridges, and badlands of diverse parent materials in open desert scrub or mountain slopes in pinyon-juniper-sagebrush woodlands, sagebrush scrub.
Elevation 10–1400 m. (0–4600 ft.) 600–3000 m. (2000–9800 ft.)
Distribution
from FNA
CA; OR
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

Rhinotropis californica occurs in western California and Oregon.

Cleistogamous and semi-cleistogamous flowers can appear earlier than chasmogamous flowers. Their flowers, fruits, and seeds are similar to those of chasmogamous flowers, but typically are smaller and without the keel beak.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rhinotropis intermontana is named for its distribution in the Intermountain region of the United States, which is bounded by the Rocky Mountains on the east, the Sierra Nevada and Cascade Range on the west, and the Mojave Desert to the south.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Rhinotropis Polygalaceae > Rhinotropis
Sibling taxa
R. acanthoclada, R. cornuta, R. heterorhyncha, R. intermontana, R. lindheimeri, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
R. acanthoclada, R. californica, R. cornuta, R. heterorhyncha, R. lindheimeri, R. maravillasensis, R. nitida, R. nudata, R. rimulicola, R. rusbyi, R. subspinosa
Synonyms Polygalacalifornica nuttall Polygalaintermontana t.
Name authority (Nuttall) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011) (T. Wendt) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 135. (2011)
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