Rhinotropis californica |
Rhinotropis |
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California milkwort |
milkwort |
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Habit | Herbs, sometimes suffrutescent, multi-stemmed, often forming a ground cover, 0.5–3.5 dm. | Herbs, perennial, subshrubs, or shrubs, single- or multi-stemmed, with or without thorns, then as modified tips of racemes. | ||||||||||||||||||||||||||||||||||||||||||||
Stems | laxly erect, decumbent, or prostrate, pubescent to subglabrous, hairs incurved. |
usually sprawling to erect, sometimes prostrate or decumbent, usually not glaucous, pubescent or glabrous. |
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Leaves | sessile or subsessile; blade ovate, elliptic, or obovate, 7–50(–60) × 3–20(–26) mm, base usually rounded to acute, sometimes cuneate, apex rounded to acute, surfaces pubescent, hairs incurved. |
alternate; sessile, subsessile, or petiolate; usually not strongly dimorphic; blade surfaces pubescent or glabrous. |
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Racemes | terminal or leaf-opposed, 1–4(–5) × 1.8–3 cm; rachis not thorn-tipped; peduncle 0–1 cm; bracts early deciduous, linear to lanceolate. |
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Inflorescences | terminal or leaf-opposed, sometimes appearing axillary if poorly developed, racemes, sometimes reduced and appearing fasciculate or aggregated into pseudopanicles; peduncle present or absent; bracts deciduous to subpersistent or persistent. |
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Pedicels | (2.5–)3.5–8.5 mm, sparsely pubescent or glabrous. |
present. |
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Flowers | usually pink, rarely white, keel distally yellow (fading white), (2.5–)9–14.5 mm, cleistogamous and semi-cleistogamous flowers mostly 2.5–5 mm, intergrading with chasmogamous flowers; sepals deciduous, elliptic, 4–6.5 mm, pubescent or glabrous; wings obovate, (7.5–)8–12 × 2.5–6 mm, glabrous or sparsely pubescent; keel (7–)8–11 mm, sac glabrous (sometimes proximally ciliate), beak oblong, (1.2–)1.6–3 × 0.7–1 mm (mostly absent in cleistogamous flowers), usually notched or contorted abaxially, rarely subentire, pubescent. |
cream, yellowish green,yellow, white, pink, rose, or purple, cleistogamous usually absent, sometimes present (in R. californica and R. lindheimeri), (2.4–)3.5–14.5 mm; sepals deciduous or persistent (when persistent, usually only upper; all persistent in R. rusbyi), sometimes appearing very slightly connate basally, pubescent or glabrous; wings deciduous, 2.5–12.5 mm, glabrous or pubescent; keel usually beaked with unlobed projection, beak sometimes reduced or obscure (rarely on all flowers unless cleistogamous, and then inflorescence usually proximal), keel glabrous or pubescent; stamens usually 7 or 8, rarely 9 (in R. acanthoclada), in chasmogamous flowers, fewer in cleistogamous flowers, not grouped; ovary 2-loculed. |
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Fruits | capsules, dehiscent, margins winged or not, glabrous or pubescent. |
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Capsules | ellipsoid to ovoid, 7.3–10.5 × 4.5–7 mm, in cleistogamous and semi-cleistogamous flowers 4.5–8 mm, base obtuse, rounded, or subtruncate, margins with narrow, entire or slightly erose wing, glabrous, margins sometimes ciliolate. |
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Seeds | 3.5–6 mm, densely pubescent; aril 1.7–4 mm, less than 1/2 length of seed. |
pubescent to subglabrous, arillate. |
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Cleistogamous | or semi-cleistogamous flowers often present terminally, on much reduced scale-leaved lateral branches from proximal (or distal) leaf axils, or terminally on leafy branches that are often leaf-opposed. |
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x | = 9. |
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2n | = 18. |
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Rhinotropis californica |
Rhinotropis |
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Phenology | Flowering spring–summer. | |||||||||||||||||||||||||||||||||||||||||||||
Habitat | Rocky or clay soils, deep duff, rich soils, serpentine soils, slopes or drainages, full sun to deep shade, open habitat, chaparral, mixed evergreen forests, oak woodlands, coniferous forests. | |||||||||||||||||||||||||||||||||||||||||||||
Elevation | 10–1400 m. (0–4600 ft.) | |||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; OR |
w United States; sc United States; Mexico; Central America (Guatemala) |
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Discussion | Rhinotropis californica occurs in western California and Oregon. Cleistogamous and semi-cleistogamous flowers can appear earlier than chasmogamous flowers. Their flowers, fruits, and seeds are similar to those of chasmogamous flowers, but typically are smaller and without the keel beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 17 (12 in the flora). Of the 17 species of Rhinotropis ranging from the southwestern United States and/or Mexico, only R. purpusii (Brandegee) J. R. Abbott extends into Guatemala. Of all the genera treated here, this is the only one that has been monographed within the last 100 years (T. L. Wendt 1978). Rhinotropis is probably sister to the Caribbean clade Phlebotaenia Grisebach, and appears to be fairly closely related also to the pantropical (although predominantly neotropical) genus Securidaca Linnaeus. Rhinotropis is largely endemic to arid regions but some species (R. californica) occur in mesic areas. The flower beak is a cylindric, conic, or contorted non-fimbriate hollow projection from the lower (or central) apex of the keel region. It is highly reduced or absent in some species. The other diagnostic features of Rhinotropis are also not monothetic across all species. Many species have the upper sepal persistent in fruit and the other sepals, including the wings (and the corolla), deciduous. Unlike other North American Polygalaceae, species of Rhinotropis often have five petals; the lateral petals are much reduced, linear, and adnate for most of their length to the staminal column; additionally, several species are shrubs and a few have thorn-tipped inflorescence axes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Polygalaceae > Rhinotropis | Polygalaceae | ||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Polygalacalifornica nuttall | Polygala section rhinotropis | ||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Nuttall) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011) | (S. F. Blake) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011) | ||||||||||||||||||||||||||||||||||||||||||||
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