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rattle-box, rattle-pod, yellow rattle

European yellow rattle

Habit Herbs, annual; hemiparasitic.
Stems

erect, not fleshy, glabrous, hairy on 2 opposite sides.

without black streaks, simple or branched, 15–40(–80) cm.

Leaves

cauline, opposite;

petiole present or absent;

blade not fleshy to +/- fleshy, not leathery, margins coarsely toothed distally.

blade lanceolate to ovate, (3.5–)5–10 mm wide, margins crenate-serrate.

Bracts

rhombic-triangular, puberulent, teeth not bristle-tipped, basal teeth scarcely longer than others.

Inflorescences

terminal, racemes, +/- secund;

bracts present.

Pedicels

present;

bracteoles absent.

Flowers

sepals 4, calyx bilaterally symmetric, flattened laterally, ovate to suborbiculate, accrescent in fruit, lobes deltate;

petals 5, corolla yellow or yellowish [bronze to bluish], compressed, strongly bilabiate, narrowly campanulate, abaxial lobes 3, adaxial 2, adaxial lip galeate, ovate, obtuse, entire, with subapical tooth on each side;

stamens 4, didynamous, filaments lanate;

staminode 0;

ovary 2-locular, placentation axile;

stigma capitate.

Corollas

15–20 mm;

tube slightly curved;

throat closed;

teeth of galea 1.5–2.5 mm.

Calyces

round-ovate in fruit, 15–20 mm, initially villous, hairs white, partly glabrescent in fruit.

Capsules

dehiscence loculicidal longitudinally.

8–12 mm.

Seeds

2–10[+], brown, kidney-shaped, flattened, wings present, absent in some R. major.

2–6, 4–5 mm.

x

= 11.

Rhinanthus

Rhinanthus major

Phenology Flowering summer.
Habitat Grasslands, clearings.
Elevation 0–100 m. (0–300 ft.)
Distribution
from USDA
North America; Europe; n Asia
[BONAP county map]
from FNA
MA; NY; Europe; Asia [Introduced in North America]
Discussion

Species ca. 50 (2 in the flora).

Although the taxonomy of Rhinanthus in the Old World is complex, the taxa found in North America are reasonably distinct. The subapical teeth on the adaxial corolla lip have been described as galea or nipples; the term teeth is used in this account. Leaf measurements are for mid-stem leaves; calyx features are for post-anthesis calyces.

The European species Rhinanthus serotinus (Schönheit ex Halácsy & H. Braun) Oborný has been reported from Maine (E. Hultén and M. Fries 1986); the authors have found no specimens to substantiate this report. If it were to be encountered in North America, it would key to R. major, from which it can be distinguished by having narrowly triangular bracts and glabrous calyces as in R. minor.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Rhinanthus alectorolophus has been applied to R. major by some authors (for example, A. Haines 2011) on the grounds that R. major is an ambiguous name (R. Soó and D. A. Webb 1972). Variants with wingless seeds are found throughout the range of R. major.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 15–20 mm; teeth of galea of corolla 1.5–2.5 mm; calyces villous, glabrescent.
R. major
1. Corollas 13–15 mm; teeth of galea of corolla (0.4–)0.6–1 mm; calyces glabrous, scabrid, or puberulent.
R. minor
Source FNA vol. 17, p. 504. Authors: Gordon C. Tucker, Bradley M. Daugherty. FNA vol. 17, p. 505.
Parent taxa Orobanchaceae Orobanchaceae > Rhinanthus
Sibling taxa
R. minor
Subordinate taxa
R. major, R. minor
Synonyms R. alectorolophus
Name authority Linnaeus: Sp. Pl. 2: 603. (1753): Gen. Pl. ed. 5, 263. (1754) Linnaeus: Amoen. Acad. 3: 53. (1756)
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