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evergreen buckthorn, holly leaf redberry, redberry

saw-tooth buckthorn, sharp-tooth buckthorn

Habit Shrubs or small trees, 1–4 m, unarmed. Shrubs or small trees, 2–3 m, usually unarmed, rarely armed with weak thorns.
Branchlets

gray, stiff, glabrous or densely and softly hirtellous.

dark purple or purple-red, glabrous.

Leaves

persistent, alternate;

petiole 2–10 mm;

blade usually dull green, sometimes reddish brown, abaxially, glossy to dull green adaxially, broadly elliptic to oblong, orbiculate, or ovate, 2–4.5 cm, distinctly coriaceous, base cuneate to nearly truncate, margins spinulose to spinose-dentate, apex rounded or obtuse to truncate or emarginate, both surfaces glabrous;

secondary veins 5–7 pairs, all diverging at nearly same angle.

deciduous, usually opposite to subopposite, sometimes alternate, usually fascicled on short shoots, separating with age;

petiole 10–28 mm;

blade green abaxially, darker green adaxially, usually broadly ovate, ovate, oblong-ovate, or ovate-cordate, sometimes ovate-orbiculate, 2.5–6(–7) cm, herbaceous, base truncate to cordate, margins spinulose-serrate, apex obtuse-rounded or acute, both surfaces glabrous or abaxial puberulent along veins;

secondary veins (3–)4–5 pairs, all diverging at nearly same angle or proximal diverging more obtusely.

Inflorescences

fascicles or flowers solitary.

fascicles or flowers solitary.

Pedicels

2–4 mm.

10–24(–30) mm.

Drupes

red, globose, 4–6(–8) mm;

stones 2.

black, globose to obovoid-globose, 7–10 mm;

stones 3.

Sepals

4.

4.

Petals

0.

4.

2n

= 24.

= 24.

Rhamnus ilicifolia

Rhamnus arguta

Phenology Flowering Jan–Jun. Flowering May–Jun.
Habitat Canyon slopes and bottoms, rock faces, open hillsides, sandstone ridges, serpentine slopes, roadsides, stream benches, riparian areas, meadows, coastal sage scrub, chaparral/desert transition, chaparral, woodlands, montane forests. Disturbed sites.
Elevation 100–2200(–2400) m. (300–7200(–7900) ft.) 200 m. (700 ft.)
Distribution
from FNA
AZ; CA; NV; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
IN; Asia (China) [Introduced in North America]
[BONAP county map]
Discussion

The only report of Rhamnus arguta as naturalized in North America is from Jasper County (F. Swink and G. Wilhelm 1994). It was identified by Swink and Wilhelm as R. arguta var. velutina Handel-Mazzetti, which is puberulent on the petioles and leaf blades (at least abaxially on veins), but considerable populational variability apparently exists within the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 48. FNA vol. 12, p. 51.
Parent taxa Rhamnaceae > Rhamnus Rhamnaceae > Rhamnus
Sibling taxa
R. alaternus, R. alnifolia, R. arguta, R. cathartica, R. crocea, R. davurica, R. japonica, R. lanceolata, R. pilosa, R. pirifolia, R. serrata, R. smithii, R. utilis
R. alaternus, R. alnifolia, R. cathartica, R. crocea, R. davurica, R. ilicifolia, R. japonica, R. lanceolata, R. pilosa, R. pirifolia, R. serrata, R. smithii, R. utilis
Synonyms R. crocea subsp. ilicifolia, R. crocea var. ilicifolia
Name authority Kellogg: Proc. Calif. Acad. Sci. 2: 37. (1863) — (as ilicifolius) Maximowicz: Mém. Acad. Imp. Sci. Saint Pétersbourg, Sér. 7, 10(11): 6, figs. 48–51. (1866)
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