Rauiella scita |
Thuidiaceae |
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rauiella moss |
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Habit | Plants small to large, usually in stiff, loose mats, dark green to golden brown. | |||||||||||||
Stem(s) | leaves erect-spreading when moist, broadly ovate to cordate-deltoid; margins loosely revolute proximally, plane distally; apex narrowly long-acuminate; laminal cells 5–6 µm, papillae 3–6. |
and branch leaves strongly differentiated; margins serrulate to crenulate ± throughout; costa single; alar cells not or poorly differentiated; laminal cells quadrate to short-rectangular, 1- or multipapillose on both surfaces or only abaxially, walls firm.; stem leaves usually spreading when moist, usually broadly ovate, concave; margins plane or recurved; apex acuminate; costa strong, ending in acumen to ± excurrent. |
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Branch leaves | appressed when dry, erect-spreading when moist, 0.4 mm; apex acute; costa pellucid proximally; laminal cells 5–6 µm, papillae 4–6; apical cell truncate, multipapillose. |
usually spreading, ovate to oblong-ovate, smaller than stem leaves; margins usually plane; apex usually gradually acuminate to obtuse; costa weaker than in stem leaves; apical laminal cell usually truncate, multipapillose, walls not porose. |
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Seta | 1–2 cm. |
reddish, elongate, smooth to roughened throughout or rarely only distally. |
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Sexual condition | autoicous or dioicous; perichaetia conspicuous, leaves erect, sometimes plicate, margins plane, apex long-acuminate, costa single, usually strong, laminal cells rectangular, more elongate than in vegetative leaves, usually smooth. |
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Capsule | with operculum obliquely short-rostrate; endostome segments narrowly perforate, cilia in groups of 2 or 3. |
usually inclined to horizontal, rarely suberect, usually asymmetric and arcuate; exothecial cell walls thin to thick, not collenchymatous; annulus differentiated; operculum usually rostrate; peristome hypnoid, double, attached at mouth; exostome teeth bordered, shouldered, external surface cross striolate basally, sometimes with overlying papillae, internal surface papillose, trabeculate; endostome basal membrane high, segments keeled, not to narrowly perforate, cilia in groups of 1–4, nodulose. |
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Calyptra | cucullate, smooth or roughened, naked. |
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Spores | 10–13 µm, finely papillose. |
spheric, usually papillose. |
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Perichaetial | leaves 1.5 mm. |
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Specialized | asexual reproduction absent. |
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Rauiella scita |
Thuidiaceae |
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Phenology | Capsules mature fall–early winter. | |||||||||||||
Habitat | Bark, base of hardwoods, mature forests | |||||||||||||
Elevation | moderate elevations | |||||||||||||
Distribution |
CT; DC; DE; IA; IN; KY; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; NS; ON; QC
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North America; Mexico; Central America; South America; West Indies; Asia; Africa |
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Discussion | Rauiella scita is more likely to be confused with Leskea (Leskeaceae) than with any other Thuidiaceae. However, the abundant paraphyllia and multipapillose laminal cells distinguish R. scita. As in R. praelonga, the distal portion of the costa of the branch leaves is covered with laminal cells, but in R. scita the apical cell of the branch leaves is truncate and multipapillose, unlike the smooth, pointed apical cell in R. praelonga. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 9, species ca. 130 (4 genera, 11 species in the flora). Thuidiaceae are characterized by pinnate branching and strongly differentiated stem and branch leaves. The stems are usually amply clothed with paraphyllia, but in smaller plants, paraphyllia are often sparse. The laminal cells of stem and branch leaves are short and papillose. Haplocladium (referred to Leskeaceae) might be confused with various Thuidiaceae. However, in Haplocladium, the paraphyllia are not papillose (as they are throughout Thuidiaceae) and branch leaves terminate in a smooth, sharp cell, rather than a truncate, multipapillose cell (except in Rauiella praelonga). Thuidiaceae were distilled by W. R. Buck and H. A. Crum (1990), emphasizing sexuality, placement of laminal cell papillae, papillosity of the cells of the paraphyllia, morphology of axillary hairs, and seta ornamentation. More recently, A. Touw (2001) provided a different generic scheme apparently based primarily on knowledge of the Asian flora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 379. | FNA vol. 28, p. 373. | ||||||||||||
Parent taxa | Thuidiaceae > Rauiella | |||||||||||||
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Synonyms | Hypnum scitum, Rauia scita, R. scita var. aestivalis, Thuidium scitum, T. scitum var. aestivale | |||||||||||||
Name authority | (P. Beauvois) Reimers: Hedwigia 76: 287. (1937) | Schimper | ||||||||||||
Web links |