Raphanus raphanistrum |
Raphanus |
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jointed charlock, jointed charlock or radish or wild-radish, wild-radish |
radish |
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Habit | Annuals, roots not fleshy; sparsely to densely pubescent. | Annuals or biennials; (roots slender or fleshy, size, shape, and color variable in cultivated forms); not scapose; glabrous or pubescent. | ||||
Stems | usually simple from base, (2–)3–8 dm, (retrorsely hispid). |
erect, unbranched or branched. |
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Leaves | basal and cauline; petiolate or subsessile; basal not rosulate, petiolate, blade margins lyrately lobed or pinnatifid to pinnatisect; cauline shortly petiolate or subsessile, blade (base not auriculate), margins dentate or lobed, (smaller and fewer-lobed than basal). |
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Basal leaves | petiole 1–6 cm; blade oblong, obovate, or oblanceolate in outline, lyrate or pinnatifid, sometimes undivided, 3–15(–22) cm × 10–50 mm, margins dentate, apex obtuse or acute; lobes 1–4 each side, oblong or ovate, to 4 cm × 20 mm (smaller than terminal). |
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Cauline leaves | (distal) subsessile; blade often undivided. |
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Racemes | (corymbose, several-flowered), usually greatly elongated in fruit. |
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Flowers | sepals 7–11 × 1–2 mm, sparsely pubescent; petals yellow or creamy white (veins dark brown or purple), 15–25 × 4–7 mm, claw to 15 mm; filaments (slender), 7–12 mm; anthers 2–2.5 mm. |
sepals erect, narrowly oblong [linear], lateral pair slightly saccate basally; petals white, creamy white, yellow, pink, or purple [lilac] (usually with darker veins), broadly obovate [suborbicular], claw differentiated from blade, (± longer than sepals, apex obtuse or emarginate [rounded]); stamens strongly tetradynamous; filaments not dilated basally; anthers oblong or oblong-linear, (apex obtuse); nectar glands (4), median pair present. |
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Fruiting pedicels | divaricate or ascending, 7–25 mm, (straight). |
divaricate, ascending, or spreading [reflexed]. |
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Fruits | cylindrical or narrowly lanceolate; valvular segment 1–1.5 mm; terminal segment (1.5–)2–11(–14) cm × (2.5–)3–8(–11) mm, (base rounded), strongly constricted between seeds (usually breaking), strongly ribbed, beak narrowly conical; style 10–50 mm. |
siliques or silicles, indehiscent, sessile, segments 2, (lomentaceous, often breaking into 1-seeded units), cylindrical, fusiform, lanceolate, or ovoid, [linear, oblong, ellipsoid], smooth or torulose to strongly moniliform, (constricted or not between seeds), terete or polygonal; (valvular segment seedless, rudimentary, or aborted, nearly as wide as pedicel; terminal segment several-seeded, corky); valves glabrous, antrorsely scabrous, or hispid; replum and septum not differentiated; ovules 2–22 per ovary; (style slender); stigma capitate, slightly 2-lobed. |
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Seeds | (reddish brown or dark brown to black), oblong or ovoid, 2.5–3.5 × 1.8–2.5 mm. |
uniseriate, plump, not winged, oblong, ovoid, or globose [subglobose]; seed coat (nearly smooth to reticulate), not mucilaginous when wetted; cotyledons conduplicate. |
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x | = 9. |
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2n | = 18. |
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Raphanus raphanistrum |
Raphanus |
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Phenology | Flowering May–Jul. | |||||
Habitat | Disturbed waste places, cultivated fields, roadsides, orchards, hill slopes | |||||
Elevation | 0-800 m (0-2600 ft) | |||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Greenland; Europe; Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Atlantic Islands, Australia]
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Eurasia [Introduced in North America; introduced also nearly worldwide] |
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Discussion | North American representatives of Raphanus raphanistrum are referable to subsp. raphanistrum. Four other subspecies are restricted to Europe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 3 (2 in the flora). Natural hybridization between Raphanus raphanistrum and R. sativus has been known since 1788, and the hybrid has been named R. ×micranthus (Uechtritz) O. E. Schulz. The transfer of some of the weedy characters from R. raphanistrum to R. sativus through natural hybridization may have played a major role in converting the latter from a crop plant into a successful weed near the coastal areas of central California (C. A. Panetsos and H. G. Baker 1968). Raphanus confusus (Greuter & Burdet) Al-Shehbaz & Warwick is known from Asia (Israel, Lebanon). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 439. | FNA vol. 7, p. 438. | ||||
Parent taxa | Brassicaceae > tribe Brassiceae > Raphanus | Brassicaceae > tribe Brassiceae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Quidproquo | |||||
Name authority | Linnaeus: Sp. Pl. 1: 669. (1753) | Linnaeus: Sp. Pl. 2: 669. (1753): Gen. Pl. ed. 5, 300. (1754) | ||||
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