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jointed charlock, jointed charlock or radish or wild-radish, wild-radish

radish

Habit Annuals, roots not fleshy; sparsely to densely pubescent. Annuals or biennials; (roots slender or fleshy, size, shape, and color variable in cultivated forms); not scapose; glabrous or pubescent.
Stems

usually simple from base, (2–)3–8 dm, (retrorsely hispid).

erect, unbranched or branched.

Leaves

basal and cauline; petiolate or subsessile;

basal not rosulate, petiolate, blade margins lyrately lobed or pinnatifid to pinnatisect;

cauline shortly petiolate or subsessile, blade (base not auriculate), margins dentate or lobed, (smaller and fewer-lobed than basal).

Basal leaves

petiole 1–6 cm;

blade oblong, obovate, or oblanceolate in outline, lyrate or pinnatifid, sometimes undivided, 3–15(–22) cm × 10–50 mm, margins dentate, apex obtuse or acute;

lobes 1–4 each side, oblong or ovate, to 4 cm × 20 mm (smaller than terminal).

Cauline leaves

(distal) subsessile;

blade often undivided.

Racemes

(corymbose, several-flowered), usually greatly elongated in fruit.

Flowers

sepals 7–11 × 1–2 mm, sparsely pubescent;

petals yellow or creamy white (veins dark brown or purple), 15–25 × 4–7 mm, claw to 15 mm;

filaments (slender), 7–12 mm;

anthers 2–2.5 mm.

sepals erect, narrowly oblong [linear], lateral pair slightly saccate basally;

petals white, creamy white, yellow, pink, or purple [lilac] (usually with darker veins), broadly obovate [suborbicular], claw differentiated from blade, (± longer than sepals, apex obtuse or emarginate [rounded]);

stamens strongly tetradynamous;

filaments not dilated basally;

anthers oblong or oblong-linear, (apex obtuse);

nectar glands (4), median pair present.

Fruiting pedicels

divaricate or ascending, 7–25 mm, (straight).

divaricate, ascending, or spreading [reflexed].

Fruits

cylindrical or narrowly lanceolate;

valvular segment 1–1.5 mm;

terminal segment (1.5–)2–11(–14) cm × (2.5–)3–8(–11) mm, (base rounded), strongly constricted between seeds (usually breaking), strongly ribbed, beak narrowly conical;

style 10–50 mm.

siliques or silicles, indehiscent, sessile, segments 2, (lomentaceous, often breaking into 1-seeded units), cylindrical, fusiform, lanceolate, or ovoid, [linear, oblong, ellipsoid], smooth or torulose to strongly moniliform, (constricted or not between seeds), terete or polygonal; (valvular segment seedless, rudimentary, or aborted, nearly as wide as pedicel; terminal segment several-seeded, corky);

valves glabrous, antrorsely scabrous, or hispid;

replum and septum not differentiated;

ovules 2–22 per ovary; (style slender);

stigma capitate, slightly 2-lobed.

Seeds

(reddish brown or dark brown to black), oblong or ovoid, 2.5–3.5 × 1.8–2.5 mm.

uniseriate, plump, not winged, oblong, ovoid, or globose [subglobose];

seed coat (nearly smooth to reticulate), not mucilaginous when wetted;

cotyledons conduplicate.

x

= 9.

2n

= 18.

Raphanus raphanistrum

Raphanus

Phenology Flowering May–Jul.
Habitat Disturbed waste places, cultivated fields, roadsides, orchards, hill slopes
Elevation 0-800 m (0-2600 ft)
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Greenland; Europe; Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Atlantic Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from USDA
Eurasia [Introduced in North America; introduced also nearly worldwide]
[BONAP county map]
Discussion

North American representatives of Raphanus raphanistrum are referable to subsp. raphanistrum. Four other subspecies are restricted to Europe.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 3 (2 in the flora).

Natural hybridization between Raphanus raphanistrum and R. sativus has been known since 1788, and the hybrid has been named R. ×micranthus (Uechtritz) O. E. Schulz. The transfer of some of the weedy characters from R. raphanistrum to R. sativus through natural hybridization may have played a major role in converting the latter from a crop plant into a successful weed near the coastal areas of central California (C. A. Panetsos and H. G. Baker 1968). Raphanus confusus (Greuter & Burdet) Al-Shehbaz & Warwick is known from Asia (Israel, Lebanon).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pale or creamy white; fruits (2.5-)3-8(-11) mm wide, strongly constricted between seeds and usually breaking, strongly ribbed, beak narrowly conical.
R. raphanistrum
1. Petals usually purple or pink, sometimes white; fruits (5-)7-13(-15) mm wide, rarely slightly constricted between seeds and usually not breaking, not ribbed, beak narrowly to broadly conical to linear.
R. sativus
Source FNA vol. 7, p. 439. FNA vol. 7, p. 438. Author: Suzanne I. Warwick.
Parent taxa Brassicaceae > tribe Brassiceae > Raphanus Brassicaceae > tribe Brassiceae
Sibling taxa
R. sativus
Subordinate taxa
R. raphanistrum, R. sativus
Synonyms Quidproquo
Name authority Linnaeus: Sp. Pl. 1: 669. (1753) Linnaeus: Sp. Pl. 2: 669. (1753): Gen. Pl. ed. 5, 300. (1754)
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