Ranunculus testiculatus |
Ranunculus subg. Ceratocephala |
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bur buttercup, hornseed buttercup, tubercled crowfoot |
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Habit | Plants villous. | |
Roots | basal, never tuberous. |
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Stems | erect or ascending, not rooting nodally, villous, not bulbous-based. |
erect, not bulbous-based, without bulbils. |
Leaves | all basal, petiolate; blade 1-2x-dissected. |
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Basal leaf | blades broadly spatulate in outline, 1-2x-dissected, 0.9-3.8 × 0.5-1.5 cm, segments linear, margins entire, apex obtuse to acuminate. |
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Inflorescences | terminal, solitary flowers. |
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Flowers | receptacle glabrous; sepals spreading, 3-6 × 1-2 mm, villous; petals yellow, 3-5 × 1-3 mm. |
sepals persistent in fruit, 5, 3-6 × 1-2 mm; petals 5, yellow; nectary scale attached basally, forming flap over nectary, glabrous, free margin entire; style present. |
Fruits | achenes, 3-locular, lateral locules empty; achene body ellipsoid, about as wide as thick, not prolonged beyond seed; wall firm, smooth; margin low narrow ridge; beak much longer than achene body. |
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Heads of achenes | cylindric, 9-16(-27) × 8-10 mm; achenes 1.6-2 × 1.8-2 mm, tomentose; beak persistent, lanceolate, 3.5-4.5 mm. |
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Tuberous | roots absent. |
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Ranunculus testiculatus |
Ranunculus subg. Ceratocephala |
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Phenology | Flowering spring (Apr–May). | |
Habitat | Disturbed areas, especially in grassland | |
Elevation | 400-2500 m (1300-8200 ft) | |
Distribution |
AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OH; OR; SD; UT; WA; WY; BC; SK; native to Eurasia [Introduced in North America]
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North America; Pacific Islands; Australia; Europe; c Asia; sw Asia; and n Africa |
Discussion | In North America, Ranunculus testiculatus seems to be expanding its range rapidly in arid and semiarid areas. A second species of this subgenus, R. falcatus Linnaeus [Ceratocephala falcata (Linnaeus) Persoon], has been reported from North America, but all reports seem to be based on misidentified material of R. testiculatus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 10 (1 in the flora). Ranunculus subg. Ceratocephalus is very different in aspect and fruit structure from other species of the genus; it is often treated as a distinct genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Ranunculaceae > Ranunculus > subg. Ceratocephala | Ranunculaceae > Ranunculus |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Ceratocephalus orthoceras, Ceratocephalus testiculatus | subg. Ceratocephalus |
Name authority | Crantz: Stirp. Austr. Fasc. 2: 97. (1763) | (Moench) L. D. Benson: Amer. J. Bot. 27: 806. (1940) |
Web links |