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bur buttercup, hornseed buttercup, tubercled crowfoot

Habit Plants villous.
Roots

basal, never tuberous.

Stems

erect or ascending, not rooting nodally, villous, not bulbous-based.

erect, not bulbous-based, without bulbils.

Leaves

all basal, petiolate;

blade 1-2x-dissected.

Basal leaf

blades broadly spatulate in outline, 1-2x-dissected, 0.9-3.8 × 0.5-1.5 cm, segments linear, margins entire, apex obtuse to acuminate.

Inflorescences

terminal, solitary flowers.

Flowers

receptacle glabrous;

sepals spreading, 3-6 × 1-2 mm, villous;

petals yellow, 3-5 × 1-3 mm.

sepals persistent in fruit, 5, 3-6 × 1-2 mm;

petals 5, yellow;

nectary scale attached basally, forming flap over nectary, glabrous, free margin entire;

style present.

Fruits

achenes, 3-locular, lateral locules empty;

achene body ellipsoid, about as wide as thick, not prolonged beyond seed;

wall firm, smooth;

margin low narrow ridge;

beak much longer than achene body.

Heads of achenes

cylindric, 9-16(-27) × 8-10 mm;

achenes 1.6-2 × 1.8-2 mm, tomentose;

beak persistent, lanceolate, 3.5-4.5 mm.

Tuberous

roots absent.

Ranunculus testiculatus

Ranunculus subg. Ceratocephala

Phenology Flowering spring (Apr–May).
Habitat Disturbed areas, especially in grassland
Elevation 400-2500 m (1300-8200 ft)
Distribution
from FNA
AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OH; OR; SD; UT; WA; WY; BC; SK; native to Eurasia [Introduced in North America]
[WildflowerSearch map]
North America; Pacific Islands; Australia; Europe; c Asia; sw Asia; and n Africa
Discussion

In North America, Ranunculus testiculatus seems to be expanding its range rapidly in arid and semiarid areas. A second species of this subgenus, R. falcatus Linnaeus [Ceratocephala falcata (Linnaeus) Persoon], has been reported from North America, but all reports seem to be based on misidentified material of R. testiculatus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 10 (1 in the flora).

Ranunculus subg. Ceratocephalus is very different in aspect and fruit structure from other species of the genus; it is often treated as a distinct genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Ranunculus > subg. Ceratocephala Ranunculaceae > Ranunculus
Sibling taxa
R. abortivus, R. acriformis, R. acris, R. adoneus, R. alismifolius, R. allegheniensis, R. allenii, R. ambigens, R. andersonii, R. aquatilis, R. arizonicus, R. arvensis, R. auricomus, R. austro-oreganus, R. bonariensis, R. bulbosus, R. californicus, R. canus, R. cardiophyllus, R. cooleyae, R. cymbalaria, R. eschscholtzii, R. fascicularis, R. fasciculatus, R. ficaria, R. flabellaris, R. flammula, R. gelidus, R. glaberrimus, R. glacialis, R. gmelinii, R. gormanii, R. harveyi, R. hebecarpus, R. hederaceus, R. hispidus, R. hydrocharoides, R. hyperboreus, R. hystriculus, R. inamoenus, R. jovis, R. kamtschaticus, R. lapponicus, R. laxicaulis, R. lobbii, R. macauleyi, R. macounii, R. macranthus, R. marginatus, R. micranthus, R. muricatus, R. nivalis, R. occidentalis, R. oresterus, R. orthorhynchus, R. pacificus, R. pallasii, R. parviflorus, R. pedatifidus, R. pensylvanicus, R. platensis, R. populago, R. pusillus, R. pygmaeus, R. ranunculinus, R. recurvatus, R. repens, R. rhomboideus, R. sabinei, R. sardous, R. sceleratus, R. sulphureus, R. trilobus, R. triternatus, R. turneri, R. uncinatus
Subordinate taxa
Synonyms Ceratocephalus orthoceras, Ceratocephalus testiculatus subg. Ceratocephalus
Name authority Crantz: Stirp. Austr. Fasc. 2: 97. (1763) (Moench) L. D. Benson: Amer. J. Bot. 27: 806. (1940)
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