Ranunculus testiculatus |
Ranunculus gmelinii |
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bur buttercup, hornseed buttercup, tubercled crowfoot |
Gmelin's buttercup, Gmelin's water buttercup, lesser yellow water crowfoot, renoncule de Gmelin, small yellow water-buttercup, yellow water crowfoot |
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Stems | erect or ascending, not rooting nodally, villous, not bulbous-based. |
prostrate or sometimes floating, glabrous or hirsute, rooting nodally. |
Leaves | basal leaves absent, cauline leaf blades reniform to circular, 3-parted, 0.6-6.5 × 1.1-9 cm, base cordate, segments again 1-3x-lobed to -dissected, margins entire or crenate, apex rounded to filiform. |
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Basal leaf | blades broadly spatulate in outline, 1-2x-dissected, 0.9-3.8 × 0.5-1.5 cm, segments linear, margins entire, apex obtuse to acuminate. |
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Flowers | receptacle glabrous; sepals spreading, 3-6 × 1-2 mm, villous; petals yellow, 3-5 × 1-3 mm. |
receptacle sparsely hispid; sepals 4-5, spreading or reflexed from base, 2-5 × 2-4 mm, glabrous or sparsely pilose; petals 4-14, 3-7 × 2-5 mm; nectary scale variable, crescent-shaped, funnel-shaped, or flaplike; style 0.2-0.4 mm. |
Heads of achenes | cylindric, 9-16(-27) × 8-10 mm; achenes 1.6-2 × 1.8-2 mm, tomentose; beak persistent, lanceolate, 3.5-4.5 mm. |
globose or ovoid, 3-8 × 3-7 mm; achenes 1-1.6 × 1-1.2 mm, glabrous; beak narrowly lanceolate or filiform, 0.4-0.8 mm. |
Tuberous | roots absent. |
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2n | = 16, 32, 64. |
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Ranunculus testiculatus |
Ranunculus gmelinii |
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Phenology | Flowering spring (Apr–May). | Flowering spring–summer (May–Sep). |
Habitat | Disturbed areas, especially in grassland | Shallow water or drying mud, wet meadows, swamps, marshes, ponds, shores of rivers |
Elevation | 400-2500 m (1300-8200 ft) | 0-2800 m (0-9200 ft) |
Distribution |
AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OH; OR; SD; UT; WA; WY; BC; SK; native to Eurasia [Introduced in North America]
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AK; CO; IA; ID; IL; ME; MI; MN; MT; ND; NM; NV; OR; UT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Eurasia
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Discussion | In North America, Ranunculus testiculatus seems to be expanding its range rapidly in arid and semiarid areas. A second species of this subgenus, R. falcatus Linnaeus [Ceratocephala falcata (Linnaeus) Persoon], has been reported from North America, but all reports seem to be based on misidentified material of R. testiculatus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ranunculus gmelinii has been divided into varieties on the basis of indument and flower size. These characters are variable and poorly correlated with one another, however, and these varieties scarcely seem natural. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Ranunculaceae > Ranunculus > subg. Ceratocephala | Ranunculaceae > Ranunculus > subg. Ranunculus > sect. Hecatonia |
Sibling taxa | ||
Synonyms | Ceratocephalus orthoceras, Ceratocephalus testiculatus | R. gmelinii subsp. purshii, R. gmelinii var. hookeri, R. gmelinii var. limosus, R. gmelinii var. prolificus, R. purshii |
Name authority | Crantz: Stirp. Austr. Fasc. 2: 97. (1763) | de Candolle: Syst. Nat. 1: 303. (1817) |
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