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glacier buttercup

bur buttercup, hornseed buttercup, tubercled crowfoot

Stems

erect or ascending from caudices, not rooting nodally, distally glabrous or brown-pilose, not bulbous-based.

erect or ascending, not rooting nodally, villous, not bulbous-based.

Basal leaf blades

reniform to broadly triangular in outline, 3-foliolate or deeply 3-parted, 1-4 × 1.6-4.4 cm, leaflets or segments 1-2x-parted, ultimate segments elliptic to oblanceolate or almost linear, margins entire or occasionally with 1-2 teeth, apex rounded to obtuse.

broadly spatulate in outline, 1-2x-dissected, 0.9-3.8 × 0.5-1.5 cm, segments linear, margins entire, apex obtuse to acuminate.

Flowers

receptacle glabrous or brown-pilose;

sepals spreading, 7-12 × 4-9 mm, brown-pilose;

petals initially white, usually becoming red with age, 9-15 × 7-14(-18) mm.

receptacle glabrous;

sepals spreading, 3-6 × 1-2 mm, villous;

petals yellow, 3-5 × 1-3 mm.

Heads of achenes

cylindric, 9-16(-27) × 8-10 mm;

achenes 1.6-2 × 1.8-2 mm, tomentose;

beak persistent, lanceolate, 3.5-4.5 mm.

Tuberous

roots absent.

roots absent.

Fruiting

heads hemispheric, 5-8 × 7-16 mm;

fruit wall smooth, not veined, loose but not inflated, winged along suture, fruits winged achenes;

achenes 2.6-3 × 1.4-2 mm, glabrous;

beak persistent, lanceolate, 0.8-2.3 mm.

Ranunculus glacialis

Ranunculus testiculatus

Phenology Flowering spring (Apr–May).
Habitat Disturbed areas, especially in grassland
Elevation 400-2500 m (1300-8200 ft)
Distribution
from FNA
AK; Eurasia
from FNA
AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OH; OR; SD; UT; WA; WY; BC; SK; native to Eurasia [Introduced in North America]
[WildflowerSearch map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In North America, Ranunculus testiculatus seems to be expanding its range rapidly in arid and semiarid areas. A second species of this subgenus, R. falcatus Linnaeus [Ceratocephala falcata (Linnaeus) Persoon], has been reported from North America, but all reports seem to be based on misidentified material of R. testiculatus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems glabrous or with few hairs at nodes; caudices well-developed; basal leaves compound, ultimate segments elliptic to oblanceolate; receptacle glabrous; Greenland.
var. glacialis
1. Stems distally brown-pilose; caudices short; basal leaves deeply parted or compound, ultimate segments oblanceolate to almost linear; receptacle brown-pilose; Alaska.
var. camissonis
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Ranunculus > subg. Crymodes Ranunculaceae > Ranunculus > subg. Ceratocephala
Sibling taxa
R. abortivus, R. acriformis, R. acris, R. adoneus, R. alismifolius, R. allegheniensis, R. allenii, R. ambigens, R. andersonii, R. aquatilis, R. arizonicus, R. arvensis, R. auricomus, R. austro-oreganus, R. bonariensis, R. bulbosus, R. californicus, R. canus, R. cardiophyllus, R. cooleyae, R. cymbalaria, R. eschscholtzii, R. fascicularis, R. fasciculatus, R. ficaria, R. flabellaris, R. flammula, R. gelidus, R. glaberrimus, R. gmelinii, R. gormanii, R. harveyi, R. hebecarpus, R. hederaceus, R. hispidus, R. hydrocharoides, R. hyperboreus, R. hystriculus, R. inamoenus, R. jovis, R. kamtschaticus, R. lapponicus, R. laxicaulis, R. lobbii, R. macauleyi, R. macounii, R. macranthus, R. marginatus, R. micranthus, R. muricatus, R. nivalis, R. occidentalis, R. oresterus, R. orthorhynchus, R. pacificus, R. pallasii, R. parviflorus, R. pedatifidus, R. pensylvanicus, R. platensis, R. populago, R. pusillus, R. pygmaeus, R. ranunculinus, R. recurvatus, R. repens, R. rhomboideus, R. sabinei, R. sardous, R. sceleratus, R. sulphureus, R. testiculatus, R. trilobus, R. triternatus, R. turneri, R. uncinatus
R. abortivus, R. acriformis, R. acris, R. adoneus, R. alismifolius, R. allegheniensis, R. allenii, R. ambigens, R. andersonii, R. aquatilis, R. arizonicus, R. arvensis, R. auricomus, R. austro-oreganus, R. bonariensis, R. bulbosus, R. californicus, R. canus, R. cardiophyllus, R. cooleyae, R. cymbalaria, R. eschscholtzii, R. fascicularis, R. fasciculatus, R. ficaria, R. flabellaris, R. flammula, R. gelidus, R. glaberrimus, R. glacialis, R. gmelinii, R. gormanii, R. harveyi, R. hebecarpus, R. hederaceus, R. hispidus, R. hydrocharoides, R. hyperboreus, R. hystriculus, R. inamoenus, R. jovis, R. kamtschaticus, R. lapponicus, R. laxicaulis, R. lobbii, R. macauleyi, R. macounii, R. macranthus, R. marginatus, R. micranthus, R. muricatus, R. nivalis, R. occidentalis, R. oresterus, R. orthorhynchus, R. pacificus, R. pallasii, R. parviflorus, R. pedatifidus, R. pensylvanicus, R. platensis, R. populago, R. pusillus, R. pygmaeus, R. ranunculinus, R. recurvatus, R. repens, R. rhomboideus, R. sabinei, R. sardous, R. sceleratus, R. sulphureus, R. trilobus, R. triternatus, R. turneri, R. uncinatus
Subordinate taxa
R. glacialis var. camissonis, R. glacialis var. glacialis
Synonyms Beckwithia glacialis Ceratocephalus orthoceras, Ceratocephalus testiculatus
Name authority Linnaeus: Sp. Pl. 1: 553. (1753) Crantz: Stirp. Austr. Fasc. 2: 97. (1763)
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