Ranunculaceae |
Delphinium |
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buttercup family, crowfoot family |
delphinium, larkspur |
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Habit | Herbs, sometimes woody or herbaceous climbers or low shrubs, perennial or annual, often rhizomatous. | Herbs, perennial, from fasciculate roots or rhizomes. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | unarmed. |
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Leaves | blade undivided or more commonly divided or compound, base cordate, sometimes truncate or cuneate, margins entire, toothed, or incised; venation pinnate or palmate. |
blade deeply palmately divided, round to pentagonal or reniform, margins entire or lobes apically crenate or lacerate, lobes of basal blades wider and fewer than those of cauline blades. |
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Inflorescences | terminal or axillary, racemes, cymes, umbels, panicles, or spikes, or flowers solitary, flowers pedicellate or sessile. |
terminal, 2-100(-more)-flowered racemes (occasionally branched, thus technically panicles), 5-40 cm or more; bracts subtending inflorescence branches; pedicels present or absent; bracteoles (on pedicels) subopposite-subalternate, not forming involucre. |
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Flowers | bisexual, sometimes unisexual, inconspicuous or showy, radially or bilaterally symmetric; sepaloid bracteoles absent; perianth hypogynous; sepals usually imbricate, 3-6(-20), distinct, often petaloid and colored, occasionally spurred; petals 0-26, distinct (connate in Consolida), plane, cup-shaped, funnel-shaped, or spurred, conspicuous or greatly reduced; nectary usually present, rarely absent; stamens 5-many, distinct; anthers dehiscing longitudinally; staminodes absent (except in Aquilegia and Clematis); pistils 1-many; styles present or absent, often persistent in fruit as beak. |
bisexual, bilaterally symmetric; sepals not persistent in fruit, 5; upper sepal 1, spurred, 8-24 mm; lateral sepals 2, ± ovate to elliptic, 8-18 mm; lower sepals 2, similar to lateral sepals; upper petals 2, spurred, enclosed in upper sepal, nectary inside tip of spur; lower petals 2, plane, ± ovate, ± 2-lobed, clawed, 2-12 mm, nectary absent; stamens 25-40; filaments with base expanded; staminodes absent between stamens and pistils; pistils 3(-5), simple; ovules 8-20 per pistil; style present. |
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Fruits | achenes, follicles, or rarely utricles, capsules, or berries, often aggregated into globose to cylindric heads. |
follicles, aggregate, sessile, ± curved-cylindric, sides prominently veined or not; beak terminal, straight, 2-4 mm. |
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Seeds | 1-many per ovary, never stalked, not arillate; endosperm abundant; embryo usually small. |
dark brown to black (often appearing white because of air in seed coat cells), rectangular to pyramidal, often ± rough surfaced. |
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x | = 8. |
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Ranunculaceae |
Delphinium |
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Distribution | Worldwide |
n temperate and arctic subtropical and; in Eastern Hemisphere; tropical mountains (s of equator in Africa) |
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Discussion | Genera ca. 60, species 1700 (22 genera, 284 species in the flora). The flowers of many species of Ranunculaceae begin to open long before anthesis, while the floral organs are just partly expanded. Only mature flowers with open anthers should be used for determination of diagnostic characteristics (especially measurements). The literature is inconsistent about the term for the whorl of organs between sepals and stamens; these may be conspicuous and petaloid, or reduced to stalked nectaries, or intermediate between the two states. They have been called petals, honey-leaves, or (when they are inconspicuous) staminodes or nectaries. We follow M. Tamura (1993) and treat as petals all organs between the sepals and stamens, except in Clematis and Aquilegia where they usually bear rudimentary anthers and clearly represent staminodes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 300 (61 in the flora). Three Eurasian species of Delphinium–D. elatum Linnaeus, D. grandiflorum Linnaeus, and D. tatsienense Franchet–have been commonly cultivated in North America. Of the nonnative taxa, only D. elatum is sporadically naturalized, as far as is known. Isolating mechanisms in Delphinium appear to be primarily ecological, geographic, and/or temporal. Where these distinctions are disrupted, introgression often exists. Hybridization occurs regularly between certain taxa, particularly in areas of disturbance (e.g., roadcuts, drainage ditches, clearcuts). The more common and easily recognized hybrids are included in the key. Many names have been misapplied in Delphinium. The few misapplied names mentioned in discussions below refer to relatively widespread problems. Unless otherwise noted, the key and descriptions refer to fresh material. Some features may be significantly altered by pressing; they can, however, usually be determined with a certain amount of effort and experience. In the descriptions, "base of cleft" refers to the point where the cleft or sinus reaches most deeply into the petal blade. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3, p. 85. | FNA vol. 3. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Jussieu | Linnaeus: Sp. Pl. 1: 530. 175: Gen. Pl. ed 5, 236. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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