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huckleberry oak

black oak

Habit Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. Trees, deciduous, to 25 m. Bark dark brown to black, deeply furrowed, ridges often broken into irregular blocks, inner bark yellow or orange.
Twigs

dark reddish brown, (1.5-)2.5-4.5(-5) mm diam., glabrous or sparsely pubescent.

Leaves

blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse;

surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs.

blade ovate to obovate, (80-)100-300 × 80-150 mm, base obtuse to truncate, inequilateral, margins with 5-9 lobes and 15-50 awns, lobes oblong or distally expanded, separated by deep sinuses, apex acute to obtuse;

surfaces abaxially pale green, glabrous except for small axillary tufts of tomentum or with scattered pubescence, especially along veins, adaxially glossy, dark green, glabrous, secondary veins raised on both surfaces.

Acorns

solitary or rarely paired;

cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent;

nut ovoid, 8-17 × 5-10 mm, apex acute;

nut scar to 3 mm diam.

biennial;

cup cup-shaped or turbinate, 7-14 mm high × 12-22 mm wide, covering 1/2 nut, cup margins not involute, outer surface puberulent, inner surface pubescent, scale tips loose, especially at margin of cup, acute to acuminate;

nut subglobose to ovoid, 10-20 × 10-18 mm, glabrate, scar diam. 5.5-12 mm.

Terminal

buds conic, 2.5 mm, scales brown with ciliate margins.

buds ovoid or ellipsoid to subconic, 6-12 mm, noticeably 5-angled in cross section, tawny- or gray-pubescent.

n

= 12 ± 1; 2n = 24.

Quercus vacciniifolia

Quercus velutina

Phenology Flowering in early summer. Flowering spring.
Habitat Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline Commonly on dry slopes and upland areas, occasionally on sandy lowlands (especially in north) and poorly drained uplands and terraces
Elevation 900-2800 m (3000-9200 ft) 0-1500 m (0-4900 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON
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[BONAP county map]
Discussion

Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The bark of this species (quercitron) is rich in tannins and was once an important source of these chemicals used for tanning leather. (The yellow dye obtained from the bark is also called quercitron.)

Native Americans used Quercus velutina medicinally for indigestion, chronic dysentery, mouth sores, chills and fevers, chapped skin, hoarseness, milky urine, lung trouble, sore eyes, and as a tonic, an antiseptic, and an emetic (D. E. Moerman 1986).

Quercus velutina reportedly hybridizes with Q. coccinea, Q. ellipsoidalis (= Q. ×paleolithicola Trelease), Q. falcata [= Q. ×willdenowiana (Dippel) Zabel] (= Q. ×pinetorum Moldenke)], Q. ilicifolia (= Q. ×rehderi Trelease), Q. imbricaria (= Q. ×leana Nuttall), Q. incana, Q. laevis, and Q. laurifolia (= Q. ×cocksii Sargent, although E. J. Palmer [1948] challenged the validity of this claim), Q. marilandica, Q. nigra, Q. palustris (= Q. ×vaga E. J. Palmer & Steyermark), Q. phellos (= Q. ×filialis Little), Q. rubra, Q. shumardii, and possibly Q. arkansana (D. M. Hunt 1989).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Protobalanus Fagaceae > Quercus > sect. Lobatae
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. tinctoria
Name authority Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) Lamarck: in J. Lamarck et al., Encycl. 1: 721. (1785)
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