Quercus vacciniifolia |
Quercus pungens |
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huckleberry oak |
pungent oak, sandpaper oak |
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Habit | Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. | Shrubs or moderate-sized trees, evergreen or subevergreen. |
Bark | light brown, papery. |
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Twigs | gray, 1-2 mm diam., short velvety-tomentose, glabrate with age. |
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Buds | dark red-brown, ca. 2 mm, sparsely pubescent. |
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Leaves | blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse; surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs. |
blade elliptic to oblong, 10-40(-90) × 10-20(-40) mm, rather thick, leathery, stiff, base rounded or minutely cordate, very rarely cuneate, margins regularly undulate-crisped, not revolute, coarsely toothed or incised with acute teeth or spinose lobes, secondary veins 5-8(-14) on each side, usually branched before passing into teeth, apex acute or obtuse, rarely rounded, spine-tipped; surfaces abaxially canescent, usually densely stellate-pubescent, and mixed with stiff, harsh, stellate hairs, often sandpapery to touch, rarely glabrate, adaxially yellowish green, glossy, usually rough and sandpapery because of minute, persistent hair bases, rarely glabrate. |
Acorns | solitary or rarely paired; cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent; nut ovoid, 8-17 × 5-10 mm, apex acute; nut scar to 3 mm diam. |
subsessile or on peduncle to 3 mm; cup shallowly to deeply cup-shaped or turbinate, to 8 mm deep × 13 mm wide, covering ca. 1/4 nut, margin thin, scales reddish brown, moderately tuberculate or keeled, densely gray-tomentose; nut light brown, broadly ovoid to subcylindric, to 10 × 10 mm, apex rounded to subacute, glabrous. |
Cotyledons | distinct. |
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Terminal | buds conic, 2.5 mm, scales brown with ciliate margins. |
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Quercus vacciniifolia |
Quercus pungens |
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Phenology | Flowering in early summer. | Flowering spring. |
Habitat | Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline | On dry limestone or igneous slopes, usually in oak, pinyon, and juniper woodlands, chaparral, and sometimes descending into desert vegetation |
Elevation | 900-2800 m (3000-9200 ft) | 800-2000 m (2600-6600 ft) |
Distribution |
CA; NV; OR
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AZ; NM; TX; Mexico (Chihuahua and Coahuila)
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Discussion | Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Numerous populations appear to be hybrid swarms between Quercus pungens and Q. vaseyana, which is sometimes treated as a variety of Q. pungens. No other evidence for a close relationship exists for these two species, and such a treatment risks erecting a polyphyletic assemblage. To the west and south within the range of Q. pungens no indication of introgression exists, and the two species are strikingly different and easily separable. I interpret the contact as secondary. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Protobalanus | Fagaceae > Quercus > sect. Quercus |
Sibling taxa | ||
Synonyms | Q. undulata var. pungens | |
Name authority | Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) | Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171. 185: not Q. pungens Gandoger 1890 |
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