Quercus vacciniifolia |
Quercus havardii |
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huckleberry oak |
Havard oak, Havard's oak, Havard's shinnery oak, sand shinnery oak, sand shinoak, shinnery oak |
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Habit | Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. | Shrubs, deciduous, low, forming clones 0.3-1.5 × 10 m, rhizomatous. |
Bark | light gray, scaly-papery. |
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Twigs | brown or grayish, 1-2.5 mm diam., glabrous or densely short grayish or yellowish tomentulose, glabrate in age. |
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Buds | dark red-brown, subglobose, ca. 2 mm, sparsely pubescent. |
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Leaves | blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse; surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs. |
blade green, often turning brownish with age, polymorphic, oblong or elliptic or sometimes lanceolate to oblanceolate or ovate to obovate, (30-)50-100 × (10-)20-50 mm, rather thick and hard, base rounded to cuneate, margins flat to revolute, at least some undulate, 2-3 rounded teeth on each side, secondary veins 5-8 on each side, much branched, apex broadly rounded, rarely acute; surfaces abaxially densely grayish or yellowish tomentulose or stellate-pubescent, sometimes only sparsely pubescent, secondary veins quite prominent, adaxially lustrous, very sparsely stellate-pubescent or glabrate, secondary veins very slightly if at all raised. |
Acorns | solitary or rarely paired; cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent; nut ovoid, 8-17 × 5-10 mm, apex acute; nut scar to 3 mm diam. |
solitary or paired, subsessile or on peduncle to 10(-18) mm; cup from deeply cup-shaped to goblet-shaped, 10-12 mm deep × 15-25 mm wide, enclosing 1/3-1/2 nut, base rounded or slightly constricted, margin very thin and smooth, scales reddish brown, triangular-ovate to long-acute, proximally moderately to markedly tuberculate, pubescent, often canescent, tips loosely appressed; nut brown, ovoid, 12-25 × 14-18 mm. |
Cotyledons | distinct. |
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Terminal | buds conic, 2.5 mm, scales brown with ciliate margins. |
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Quercus vacciniifolia |
Quercus havardii |
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Phenology | Flowering in early summer. | Flowering spring. |
Habitat | Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline | Deep, shifting or stabilized sand dunes, off deep sands in putative hybrid populations |
Elevation | 900-2800 m (3000-9200 ft) | 500-1500 m (1600-4900 ft) |
Distribution |
CA; NV; OR
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NM; OK; TX
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Discussion | Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Individual clones emerging to heights of 2-3 m from thickets occur sporadically across the Texas range of Quercus havardii and express some characteristics of Q. stellata, such as more deeply lobed leaves and smaller acorns. Such putative hybrids increase in frequency in the eastern part of the range of the species. Material of Quercus havardii from the Navajo Basin of Utah and adjacent Arizona has been treated as Q. havardii var. tuckeri Welsh. Welsh followed J. M. Tucker (1970) and interpreted these intermediate populations as putative hybrids between Q. havardii and both Q. turbinella and Q. gambelii. Giving varietal rank, instead of nothospecies status, to such populations seems arbitrary, and it certainly is inconsistent with their putative hybrid origins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Protobalanus | Fagaceae > Quercus > sect. Quercus |
Sibling taxa | ||
Name authority | Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) | Rydberg: Bull. New York Bot. Gard. 2: 213. (1901) |
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