Flora of North America species comparison

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Quercus vacciniifolia

Quercus cornelius-mulleri

huckleberry oak

desert scrub oak, muller oak, muller's oak
Habit Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. Shrubs, evergreen or subevergreen, densely branched, 1-2(-3) m. Bark gray, scaly.
Twigs

gray, yellowish, or brownish, 1-1.5 mm diam., densely tomentulose, rarely glabrate.
Buds

dull brown, ovoid, 2.5-3 mm, glabrous except for margins of scales.
Leaves

blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse;

surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs.

blade strongly bicolored, ovate to oblong or narrowly obovate, 15-35 ×10-20 mm, rather thick and leathery, base cuneate or attenuate-rounded, margins entire or irregularly, shallowly toothed, teeth mucronate, rarely spinose, secondary veins 6-7 on each side, apex rounded or acute;

surfaces abaxially whitish, densely covered with minute, compact, appressed, (8-)10-14(-16)-rayed stellate hairs less than 0.2 mm diam. (lateral fusion of rays visible under high magnification), without glandular seriate hairs, adaxially dull, light green, grayish, or yellowish green, with scattered, appressed-stellate hairs to 0.2 mm diam.
Acorns

solitary or rarely paired;

cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent;

nut ovoid, 8-17 × 5-10 mm, apex acute;

nut scar to 3 mm diam.

solitary or clustered, subsessile;

cup deeply cup-shaped or turbinate, to 5-13 mm deep × 12-20 mm wide, scales whitish or cream, strongly tuberculate especially near base of cup;

nut dark brown, fusiform or cylyndric, 20-30 × 10-30 mm.
Cotyledons

distinct.
Terminal

buds conic, 2.5 mm, scales brown with ciliate margins.

Quercus vacciniifolia

Quercus cornelius-mulleri
Phenology Flowering in early summer. Flowering early spring.
Habitat Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline Open chaparral, pinyon and juniper woodlands, desert margins, often on loose granitic soils
Elevation 900-2800 m (3000-9200 ft) 1000-1800 m (3300-5900 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus cornelius-mulleri is easily distinguished from other California scrub oaks by the strongly bicolored leaves, dense minute tomentum of the abaxial leaf surface, and large acute acorns in deep tuberculate cups. Of greater difficulty are swarms of putative hybrids with Q. engelmannii, sometimes referred to as Q. ×acutidens.

In Joshua Tree National Monument a lone tree and several shrubs appear to be hybrids and backcrosses between Quercus cornelius-mulleri and Q. lobata. This tree is the basis of Quercus ×munzii J. M. Tucker.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Protobalanus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Name authority Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) Nixon & K. P. Steele: Madroño 28: 210. (1981)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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