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huckleberry oak

chêne bicolore, swamp white oak

Habit Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. Trees, deciduous, to 30 m. Bark dark gray, scaly or flat-ridged.
Twigs

light brown or tan, 2-3(-4) mm diam., glabrous.

Buds

light or dark brown, globose to ovoid, 2-3 mm, glabrous.

Leaves

blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse;

surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs.

blade obovate to narrowly elliptic or narrowly obovate, (79-)120-180(-215) × (40-)70-110(-160) mm, base narrowly cuneate to acute, margins regularly toothed, or entire with teeth in distal 1/2 only, or moderately to deeply lobed, or sometimes lobed proximally and toothed distally, secondary veins arched, divergent, (3-)5-7 on each side, apex broadly rounded or ovate;

surfaces abaxially light green or whitish, with minute, flat, appressed-stellate hairs and erect, 1-4-rayed hairs, velvety to touch, adaxially dark green, glossy, glabrous.

Acorns

solitary or rarely paired;

cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent;

nut ovoid, 8-17 × 5-10 mm, apex acute;

nut scar to 3 mm diam.

1-3(-5) mm, on thin axillary peduncle (20-)40-70 mm;

cup hemispheric or turbinate, 10-15 mm deep × 15-25 mm wide, enclosing 1/2-3/4 nut, scales closely appressed, finely grayish tomentose, those near rim of cup often with short, stout, irregularly recurved and sometimes branched, spinose awns emerging from tubercle;

nut light brown, ovoid-ellipsoid or oblong, (12-)15-21(-25) × 9-18 mm, glabrous.

Cotyledons

distinct.

Terminal

buds conic, 2.5 mm, scales brown with ciliate margins.

2n

= 24.

Quercus vacciniifolia

Quercus bicolor

Phenology Flowering in early summer. Flowering in spring.
Habitat Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline Low swamp forests, moist slopes, poorly drained uplands
Elevation 900-2800 m (3000-9200 ft) 0-1000 m (0-3300 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DE; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Putative hybrids between Quercus bicolor and Q. macrocarpa are common in areas of contact. The hybrids tend to have more deeply lobed leaves and varying degrees of development of awns as a fringe along the margin of the acorn cup. Such characteristics occur sporadically throughout many populations of Q. bicolor; in some cases they may occur because of subtle introgression.

The Iroquois used Quercus bicolor in the treatment of cholera, broken bones, consumption, and as a witchcraft medicine (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Protobalanus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. bicolor var. angustifolia, Q. bicolor var. cuneiformis, Q. bicolor var. platanoides, Q. platanoides
Name authority Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) Willdenow: in G. H. E. Muhlenberg, Ges. Naturf. Freunde Berlin Neue Schriften 3: 396. (1801)
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