FNA: Quercus tomentella vs. Quercus lobata

	Quercus tomentella	Quercus lobata
	Channel Island oak, island live oak, island oak	California white oak, roble, valley oak
Habit	Trees, to 20 m. Twigs branching at 45° angles, reddish brown, 3-4 mm diam., somewhat rigid, densely tomentose, persistent into 2d year.	Trees, deciduous, to 25(-35) m, usually with solitary trunks.
Bark		gray, scaly, deeply checkered in age.
Twigs		yellowish, gray, occasionally reddish, 2-4 mm diam., densely or sparsely tomentulose.
Buds		yellowish or light brown, ovoid, (2-)3-5(-6) mm, apex occasionally acute, densely pubescent.
Leaves	blade wavy or distinctly concave, oblong-lanceolate or elliptic, acuminate, (30-)70-10(-120) × 25-40 mm, leathery and brittle, base obtuse to cordate, secondary veins 8-10(-12) pairs, branching at 45-50° angles, strongly pinnate, raised abaxially, often sunken adaxially, margins often strongly revolute, with slightly thickened cell walls, entire or crenate to dentate with mucronate teeth, apex rounded or acute, mucronate; surfaces abaxially densely tomentose with whitish nonglandular hairs, midrib pilose, adaxially glossy dark green, sparsely pubescent.	blade broadly obovate or elliptic, moderately to deeply lobed, (40-)50-100(-120) × 30-60(-75) mm, base rounded-attenuate, cuneate, or truncate, rarely subcordate, margins with sinuses usually reaching more than 1/2 distance to midrib, lobes oblong or spatulate, obtuse, rounded, or blunt, secondary veins 5-10 on each side, apex broadly rounded; surfaces abaxially whitish or light green, densely to sparsely covered with interlocking appressed or semi-erect, 8-10(-14)-rayed stellate hairs, adaxially dark green or grayish, glossy or somewhat scurfy because of sparse stellate hairs.
Acorns	solitary or rarely paired; cup shallowly cup-shaped, 4-8 mm deep × 15-30 mm wide, scales laterally connate, appressed, deeply imbedded in tomentum, with only thin, brown, elongated apices visible, tuberculate, densely whitish brown tomentose throughout; nut ovoid, 20-30 × 15-20 mm, apex rounded.	solitary or paired, subsessile; cup deeply cup-shaped, hemispheric or turbinate, rim thick, 10-30 mm deep × 14-30 mm wide, scales grayish or cream, more acute near rim, strongly and irregularly tuberculate, especially toward base of cup; nut light brown, oblong or fusiform, 30-60 × (12-)15-25 mm, tapering to acute or rounded apex.
Cotyledons		distinct.
Terminal	buds conic, 7-10 mm, scales brown with ciliate margins.

	Quercus tomentella	Quercus lobata
Phenology	Flowering in spring, occasionally in fall.	Flowering late winter–early spring.
Habitat	Lower portions of steep canyons and occasionally ridge tops	Valley floors and moderate slopes, open grasslands, savannah and oak woodlands, riparian areas in chaparral
Elevation	100-650 m (300-2100 ft)	0-1700 m (0-5600 ft)
Distribution	CA; Mexico (Baja California and on Guadalupe Island) [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	The insular endemic Quercus tomentella is a relict as evidenced by its widespread representation in mainland late Tertiary fossil floras. Hybridization with Q. chrysolepis is apparent on the Channel Islands: Santa Catalina, Santa Cruz, and possibly San Clemente and Anacapa. Putative hybrids have been observed in narrow zones of contact on the islands of Santa Cruz and Santa Catalina. On those islands, taxonomically distinct individuals of Q. chrysolepis occur at the highest elevations, whereas Q. tomentella generally is found in moist canyons at lower elevations. Populations of Channel Island oak are in decline because of overgrazing and poor seedling recruitment. The greatest number of populations occur on Santa Rosa Island, and those are taxonomically and genetically noteworthy because Q. chrysolepis apparently is absent from the island. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Mature trees of Quercus lobata are among the largest oaks of the United States. The species hybridizes with numerous other species, but the hybrids are not common in most parts of its range. On Santa Cruz and Santa Catalina islands, however, occur extensive and relatively stable populations that show intermediate characteristics with Q. pacifica (see treatment). The hybrids have been given the name Q. ×macdonaldii, and they differ from Q. lobata in the following: leaf sinuses reaching less than half the distance to the midrib; leaves usually smaller, the lobes often more acute and brighter green; and acorns smaller, with more acute apices. The Yuri used Quercus lobata in the treatment of diarrhea (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Protobalanus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms		Q. hindsii, Q. lobata var. hindsii, Q. longiglanda
Name authority	Engelmann: Trans. Acad. Sci. St. Louis, 3: 393. (1877)	Nee.: Anales Ci. Nat. 3: 277. (1801)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus lobata

Channel Island oak, island live oak, island oak

California white oak, roble, valley oak

Habit

Trees, to 20 m. Twigs branching at 45° angles, reddish brown, 3-4 mm diam., somewhat rigid, densely tomentose, persistent into 2d year.

Trees, deciduous, to 25(-35) m, usually with solitary trunks.

Bark

gray, scaly, deeply checkered in age.

Twigs

yellowish, gray, occasionally reddish, 2-4 mm diam., densely or sparsely tomentulose.

Buds

yellowish or light brown, ovoid, (2-)3-5(-6) mm, apex occasionally acute, densely pubescent.

Leaves

blade wavy or distinctly concave, oblong-lanceolate or elliptic, acuminate, (30-)70-10(-120) × 25-40 mm, leathery and brittle, base obtuse to cordate, secondary veins 8-10(-12) pairs, branching at 45-50° angles, strongly pinnate, raised abaxially, often sunken adaxially, margins often strongly revolute, with slightly thickened cell walls, entire or crenate to dentate with mucronate teeth, apex rounded or acute, mucronate;

surfaces abaxially densely tomentose with whitish nonglandular hairs, midrib pilose, adaxially glossy dark green, sparsely pubescent.

blade broadly obovate or elliptic, moderately to deeply lobed, (40-)50-100(-120) × 30-60(-75) mm, base rounded-attenuate, cuneate, or truncate, rarely subcordate, margins with sinuses usually reaching more than 1/2 distance to midrib, lobes oblong or spatulate, obtuse, rounded, or blunt, secondary veins 5-10 on each side, apex broadly rounded;

surfaces abaxially whitish or light green, densely to sparsely covered with interlocking appressed or semi-erect, 8-10(-14)-rayed stellate hairs, adaxially dark green or grayish, glossy or somewhat scurfy because of sparse stellate hairs.

Acorns

solitary or rarely paired;

cup shallowly cup-shaped, 4-8 mm deep × 15-30 mm wide, scales laterally connate, appressed, deeply imbedded in tomentum, with only thin, brown, elongated apices visible, tuberculate, densely whitish brown tomentose throughout;

nut ovoid, 20-30 × 15-20 mm, apex rounded.

solitary or paired, subsessile;

cup deeply cup-shaped, hemispheric or turbinate, rim thick, 10-30 mm deep × 14-30 mm wide, scales grayish or cream, more acute near rim, strongly and irregularly tuberculate, especially toward base of cup;

nut light brown, oblong or fusiform, 30-60 × (12-)15-25 mm, tapering to acute or rounded apex.

Cotyledons

distinct.

Terminal

buds conic, 7-10 mm, scales brown with ciliate margins.

Quercus tomentella

Quercus lobata

Phenology

Flowering in spring, occasionally in fall.

Flowering late winter–early spring.

Habitat

Lower portions of steep canyons and occasionally ridge tops

Valley floors and moderate slopes, open grasslands, savannah and oak woodlands, riparian areas in chaparral

Elevation

100-650 m (300-2100 ft)

0-1700 m (0-5600 ft)

Distribution

CA; Mexico (Baja California and on Guadalupe Island)

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

The insular endemic Quercus tomentella is a relict as evidenced by its widespread representation in mainland late Tertiary fossil floras. Hybridization with Q. chrysolepis is apparent on the Channel Islands: Santa Catalina, Santa Cruz, and possibly San Clemente and Anacapa. Putative hybrids have been observed in narrow zones of contact on the islands of Santa Cruz and Santa Catalina. On those islands, taxonomically distinct individuals of Q. chrysolepis occur at the highest elevations, whereas Q. tomentella generally is found in moist canyons at lower elevations. Populations of Channel Island oak are in decline because of overgrazing and poor seedling recruitment. The greatest number of populations occur on Santa Rosa Island, and those are taxonomically and genetically noteworthy because Q. chrysolepis apparently is absent from the island.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mature trees of Quercus lobata are among the largest oaks of the United States. The species hybridizes with numerous other species, but the hybrids are not common in most parts of its range. On Santa Cruz and Santa Catalina islands, however, occur extensive and relatively stable populations that show intermediate characteristics with Q. pacifica (see treatment). The hybrids have been given the name Q. ×macdonaldii, and they differ from Q. lobata in the following: leaf sinuses reaching less than half the distance to the midrib; leaves usually smaller, the lobes often more acute and brighter green; and acorns smaller, with more acute apices.

The Yuri used Quercus lobata in the treatment of diarrhea (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Protobalanus

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. hindsii, Q. lobata var. hindsii, Q. longiglanda

Name authority

Engelmann: Trans. Acad. Sci. St. Louis, 3: 393. (1877)

Nee.: Anales Ci. Nat. 3: 277. (1801)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus tomentella

Quercus lobata

Quercus tomentella

Quercus lobata