FNA: Quercus tomentella vs. Quercus laceyi

	Quercus tomentella	Quercus laceyi
	Channel Island oak, island live oak, island oak	lacey oak
Habit	Trees, to 20 m. Twigs branching at 45° angles, reddish brown, 3-4 mm diam., somewhat rigid, densely tomentose, persistent into 2d year.	Trees, deciduous, to 5-8(-10) m. Bark light colored, papery or scaly.
Twigs		gray, 1.5-2 mm diam., pubescent with erect stellate hairs, these soon shed, at maturity reddish and pruinose to tan and glabrous.
Buds		brown, ovoid to ovoid-lanceoloid, 1.5-3 × 1-2 mm, apex acute, glabrous.
Leaves	blade wavy or distinctly concave, oblong-lanceolate or elliptic, acuminate, (30-)70-10(-120) × 25-40 mm, leathery and brittle, base obtuse to cordate, secondary veins 8-10(-12) pairs, branching at 45-50° angles, strongly pinnate, raised abaxially, often sunken adaxially, margins often strongly revolute, with slightly thickened cell walls, entire or crenate to dentate with mucronate teeth, apex rounded or acute, mucronate; surfaces abaxially densely tomentose with whitish nonglandular hairs, midrib pilose, adaxially glossy dark green, sparsely pubescent.	blade blue-green, glaucous, obovate or elliptic, (20-)40-90(-210) × (20-)30-60(-110) mm, thin, base cuneate and decurrent on petiole to rounded or rarely somewhat cordate, margins thin, flat, entire to shallowly lobed or (rarely in shade forms) deeply lobed, lobes if present oblong, squarish, often retuse, secondary veins 6-9 on each side, each terminating in tooth or arching near margins, apex broadly rounded, retuse; surfaces abaxially whitish, with erect stellate hairs, hairs shed as leaves expand, becoming glabrous, glaucous, adaxially glabrous, glaucous.
Acorns	solitary or rarely paired; cup shallowly cup-shaped, 4-8 mm deep × 15-30 mm wide, scales laterally connate, appressed, deeply imbedded in tomentum, with only thin, brown, elongated apices visible, tuberculate, densely whitish brown tomentose throughout; nut ovoid, 20-30 × 15-20 mm, apex rounded.	annual, solitary or paired, subsessile or on short peduncle to 10(-20) mm in leaf axil; cup saucer-shaped or shallowly cup-shaped, 4-7 mm deep × 10-12(-18) mm wide, enclosing 1/3 nut or less, scales moderately tuberculate, finely tomentose; nut oblong or barrel-shaped, often flattened at both ends, (11-)13-15(-20) × 9-11(-14) mm.
Cotyledons		distinct.
Terminal	buds conic, 7-10 mm, scales brown with ciliate margins.

	Quercus tomentella	Quercus laceyi
Phenology	Flowering in spring, occasionally in fall.	Flowering in spring.
Habitat	Lower portions of steep canyons and occasionally ridge tops	Limestone hills, woodlands and riparian forests, canyons and streamsides
Elevation	100-650 m (300-2100 ft)	350-2200 m (1100-7200 ft)
Distribution	CA; Mexico (Baja California and on Guadalupe Island) [WildflowerSearch map] [BONAP county map]	TX; Mexico (Coahuila and Nuevo León) [BONAP county map]
Discussion	The insular endemic Quercus tomentella is a relict as evidenced by its widespread representation in mainland late Tertiary fossil floras. Hybridization with Q. chrysolepis is apparent on the Channel Islands: Santa Catalina, Santa Cruz, and possibly San Clemente and Anacapa. Putative hybrids have been observed in narrow zones of contact on the islands of Santa Cruz and Santa Catalina. On those islands, taxonomically distinct individuals of Q. chrysolepis occur at the highest elevations, whereas Q. tomentella generally is found in moist canyons at lower elevations. Populations of Channel Island oak are in decline because of overgrazing and poor seedling recruitment. The greatest number of populations occur on Santa Rosa Island, and those are taxonomically and genetically noteworthy because Q. chrysolepis apparently is absent from the island. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Material from Texas and northeastern Mexico, excluding the type, has been incorrectly referred to Quercus glaucoides M. Martens & Galeotti by some authors (K. C. Nixon and C. H. Muller 1992). On the Edwards Plateau of Texas, Quercus laceyi occurs mostly at 350-600 m elevation; in Coahuila and Nuevo León, it occurs at 1500-2200 m. This species is sometimes associated with remnant mesic forests, which include Acer grandidentatum Nuttall, Tilia species, Quercus muhlenbergii Engelmann, and various pine and other oak species. The leaves are shallowly lobed or entire, although occasional specimens on moist sites are deeply lobed and resemble the leaves of Q. alba in outline. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Protobalanus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms		Q. breviloba subsp. laceyi
Name authority	Engelmann: Trans. Acad. Sci. St. Louis, 3: 393. (1877)	Small: Bull. Torrey Bot. Club 28: 358. (1901)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus laceyi

Channel Island oak, island live oak, island oak

lacey oak

Habit

Trees, to 20 m. Twigs branching at 45° angles, reddish brown, 3-4 mm diam., somewhat rigid, densely tomentose, persistent into 2d year.

Trees, deciduous, to 5-8(-10) m. Bark light colored, papery or scaly.

Twigs

gray, 1.5-2 mm diam., pubescent with erect stellate hairs, these soon shed, at maturity reddish and pruinose to tan and glabrous.

Buds

brown, ovoid to ovoid-lanceoloid, 1.5-3 × 1-2 mm, apex acute, glabrous.

Leaves

blade wavy or distinctly concave, oblong-lanceolate or elliptic, acuminate, (30-)70-10(-120) × 25-40 mm, leathery and brittle, base obtuse to cordate, secondary veins 8-10(-12) pairs, branching at 45-50° angles, strongly pinnate, raised abaxially, often sunken adaxially, margins often strongly revolute, with slightly thickened cell walls, entire or crenate to dentate with mucronate teeth, apex rounded or acute, mucronate;

surfaces abaxially densely tomentose with whitish nonglandular hairs, midrib pilose, adaxially glossy dark green, sparsely pubescent.

blade blue-green, glaucous, obovate or elliptic, (20-)40-90(-210) × (20-)30-60(-110) mm, thin, base cuneate and decurrent on petiole to rounded or rarely somewhat cordate, margins thin, flat, entire to shallowly lobed or (rarely in shade forms) deeply lobed, lobes if present oblong, squarish, often retuse, secondary veins 6-9 on each side, each terminating in tooth or arching near margins, apex broadly rounded, retuse;

surfaces abaxially whitish, with erect stellate hairs, hairs shed as leaves expand, becoming glabrous, glaucous, adaxially glabrous, glaucous.

Acorns

solitary or rarely paired;

cup shallowly cup-shaped, 4-8 mm deep × 15-30 mm wide, scales laterally connate, appressed, deeply imbedded in tomentum, with only thin, brown, elongated apices visible, tuberculate, densely whitish brown tomentose throughout;

nut ovoid, 20-30 × 15-20 mm, apex rounded.

annual, solitary or paired, subsessile or on short peduncle to 10(-20) mm in leaf axil;

cup saucer-shaped or shallowly cup-shaped, 4-7 mm deep × 10-12(-18) mm wide, enclosing 1/3 nut or less, scales moderately tuberculate, finely tomentose;

nut oblong or barrel-shaped, often flattened at both ends, (11-)13-15(-20) × 9-11(-14) mm.

Cotyledons

distinct.

Terminal

buds conic, 7-10 mm, scales brown with ciliate margins.

Quercus tomentella

Quercus laceyi

Phenology

Flowering in spring, occasionally in fall.

Flowering in spring.

Habitat

Lower portions of steep canyons and occasionally ridge tops

Limestone hills, woodlands and riparian forests, canyons and streamsides

Elevation

100-650 m (300-2100 ft)

350-2200 m (1100-7200 ft)

Distribution

CA; Mexico (Baja California and on Guadalupe Island)

[WildflowerSearch map]

[BONAP county map]

TX; Mexico (Coahuila and Nuevo León)

[BONAP county map]

Discussion

The insular endemic Quercus tomentella is a relict as evidenced by its widespread representation in mainland late Tertiary fossil floras. Hybridization with Q. chrysolepis is apparent on the Channel Islands: Santa Catalina, Santa Cruz, and possibly San Clemente and Anacapa. Putative hybrids have been observed in narrow zones of contact on the islands of Santa Cruz and Santa Catalina. On those islands, taxonomically distinct individuals of Q. chrysolepis occur at the highest elevations, whereas Q. tomentella generally is found in moist canyons at lower elevations. Populations of Channel Island oak are in decline because of overgrazing and poor seedling recruitment. The greatest number of populations occur on Santa Rosa Island, and those are taxonomically and genetically noteworthy because Q. chrysolepis apparently is absent from the island.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Material from Texas and northeastern Mexico, excluding the type, has been incorrectly referred to Quercus glaucoides M. Martens & Galeotti by some authors (K. C. Nixon and C. H. Muller 1992).

On the Edwards Plateau of Texas, Quercus laceyi occurs mostly at 350-600 m elevation; in Coahuila and Nuevo León, it occurs at 1500-2200 m. This species is sometimes associated with remnant mesic forests, which include Acer grandidentatum Nuttall, Tilia species, Quercus muhlenbergii Engelmann, and various pine and other oak species. The leaves are shallowly lobed or entire, although occasional specimens on moist sites are deeply lobed and resemble the leaves of Q. alba in outline.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Protobalanus

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. breviloba subsp. laceyi

Name authority

Engelmann: Trans. Acad. Sci. St. Louis, 3: 393. (1877)

Small: Bull. Torrey Bot. Club 28: 358. (1901)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus tomentella

Quercus laceyi

Quercus tomentella

Quercus laceyi