FNA: Quercus texana vs. Quercus wislizenii

	Quercus texana	Quercus wislizenii
	Nuttall's oak, Texas red oak	interior live oak, Sierra live oak
Habit	Trees, deciduous, to 25 m. Bark dark brown with flat ridges divided by shallow fissures.	Trees or shrubs, evergreen, to 22 m. Bark nearly black, deeply furrowed with broad scaly ridges.
Twigs	red-brown to gray, 1.5-3(-3.5) mm diam., glabrous.	brown to red-brown, 1.5-3 mm diam., glabrous or sparsely pubescent.
Leaves	blade ovate to elliptic or obovate, 75-200 × 55-130 mm, base cuneate to almost truncate, often inequilateral, margins with 6-11 lobes and 9-24 awns, lobes oblong to distally expanded, rarely falcate, apex acute; surfaces abaxially glabrous except for conspicuous axillary tufts of tomentum, veins raised, adaxially planar, glabrous.	blade circular to oblong, usually ovate, planar, 25-70 × 20-50 mm, base obtuse to cordate, margins entire or spinose with up to 16 awns, apex acute to rounded; surfaces abaxially and adaxially glabrous, veins little raised on either surface.
Acorns	biennial; cup thin (scale bases visible on inner surface), deeply goblet-shaped with pronounced constriction at base, 10-16 mm high × 15-22 mm wide, covering 1/3-1/2 nut, outer surface glabrous to sparsely puberulent, inner surface sparsely to uniformly pubescent, scale tips appressed, acute; nut broadly ovoid to broadly ellipsoid, 15-26 × 13-18 mm, glabrous or sparsely puberulent, scar diam. 8-13 mm, scar often orangish.	biennial; cup deeply and narrowly cup-shaped or U-shaped, 9-19 mm high × 7-18 mm wide, covering 1/3-1/2(-2/3) nut, outer surface glabrous to sparsely puberulent, inner surface glabrous or pubescent on innermost 1/3, occasionally uniformly pubescent, scales acute, tips loose; nut narrowly conic or ovoid to narrowly oblong, 21-44 × 8-14 mm, glabrous, scar diam. 2.5-7.5 mm.
Terminal	buds gray to gray-brown, ovoid, 3-7 mm, glabrous or with scales somewhat ciliate at apex.	buds light chestnut brown to dark reddish brown, ovoid to conic, 3-9 mm, glabrous or with tuft of minute hairs at apex.
2n		= 24.

	Quercus texana	Quercus wislizenii
Phenology	Flowering spring.	Flowering late spring.
Habitat	Flood plains and bottomlands	Valleys, slopes, and sand chaparral
Elevation	0-200 m (0-700 ft)	300-1900 m (1000-6200 ft)
Distribution	AL; AR; IL; KY; LA; MO; MS; TN; TX [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map]
Discussion	For many years the name Quercus texana was erroneously used for Q. buckleyi (L. J. Dorr and K. C. Nixon 1985). A few authors have also used the name for Q. gravesii. Quercus nuttallii E. J. Palmer var. cachensis E. J. Palmer was described as a small-fruited form (nuts 16-18 × 12-16 mm) from specimens collected in east-central Arkansas (E. J. Palmer 1937). Noting the similarity between Q. nuttallii var. cachensis and Q. palustris, Palmer discounted the possibility of the former being of hybrid origin because (1) he had not observed Q. palustris in the type locality, and (2) the leaves and buds of the former were essentially the same as in Q. nuttallii var. nuttallii. C. H. Muller (1942), on the other hand, argued that Quercus nuttallii was nothing more than a form [forma nuttallii (E. J. Palmer) C. H. Muller] of Q. palustris. This is a puzzling conclusion because it was based largely on the premise that Q. nuttallii occurred "...with the parent species throughout a large part of the latter's southern range (Mississippi to eastern Texas and southeastern Missouri)." The range of Q. palustris does not extend into Mississippi or eastern Texas, although its range does overlap that of Q. texana in eastern Arkansas and southeastern Missouri. E. J. Palmer (1948) and D. M. Hunt (1989) have suggested hybridization with Q. shumardii and Q. nigra, respectively. See L. J. Dorr and K. C. Nixon (1985) for an explanation of the nomenclatural confusion regarding this taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Shrubs with oval leaves 25-38 mm and margins entire or deeply lobed-dentate may be treated as Quercus wislizenii var. frutescens. J. M. Tucker (1993) treated Q. parvula as a distinct species, distinguished from Q. wislizenii by its larger leaves (30-90 versus 20-50 mm), by the dull, olive-green, abaxial leaf surface (versus shiny, yellow-green), and by nuts that are abruptly tapered proximal to the middle (versus gradually tapered). Tucker recognized two varieties of Quercus parvula: Q. parvula var. parvula is a shrub of 1-3 m and Q. parvula var. shrevei is a tree less than 17 m. S. K. Langer (1993) recognized a third variety, Q. parvula var. tamalpaisensis,based on several small populations on or near Mount Tamalpais, differentiated primarily by having larger leaves (50-160 × 20-60 mm) with attenuate-dentate margins. Quercus wislizenii reportedly hybridizes with Q. agrifolia and Q. kelloggii (W. B. Brophy and D. R. Parnell 1974). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Lobatae	Fagaceae > Quercus > sect. Lobatae
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana
Synonyms	Q. nuttallii, Q. rubra var. texana, Q. shumardii var. texana	Q. parvula, Q. parvula var. shrevei, Q. parvula var. tamalpaisensis, Q. wislizenii var. frutescens
Name authority	Buckley: Proc. Acad. Nat. Sci. Philadelphia 12: 444. (1860)	A. de Candolle: in A. P. de Candolle and A. L. P. de Candolle, Prodr. 16(2): 67. (1864)
Web links	Local Web sites: IL Wildflowers, LA Plants, MO Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus wislizenii

Nuttall's oak, Texas red oak

interior live oak, Sierra live oak

Habit

Trees, deciduous, to 25 m. Bark dark brown with flat ridges divided by shallow fissures.

Trees or shrubs, evergreen, to 22 m. Bark nearly black, deeply furrowed with broad scaly ridges.

Twigs

red-brown to gray, 1.5-3(-3.5) mm diam., glabrous.

brown to red-brown, 1.5-3 mm diam., glabrous or sparsely pubescent.

Leaves

blade ovate to elliptic or obovate, 75-200 × 55-130 mm, base cuneate to almost truncate, often inequilateral, margins with 6-11 lobes and 9-24 awns, lobes oblong to distally expanded, rarely falcate, apex acute;

surfaces abaxially glabrous except for conspicuous axillary tufts of tomentum, veins raised, adaxially planar, glabrous.

blade circular to oblong, usually ovate, planar, 25-70 × 20-50 mm, base obtuse to cordate, margins entire or spinose with up to 16 awns, apex acute to rounded;

surfaces abaxially and adaxially glabrous, veins little raised on either surface.

Acorns

biennial;

cup thin (scale bases visible on inner surface), deeply goblet-shaped with pronounced constriction at base, 10-16 mm high × 15-22 mm wide, covering 1/3-1/2 nut, outer surface glabrous to sparsely puberulent, inner surface sparsely to uniformly pubescent, scale tips appressed, acute;

nut broadly ovoid to broadly ellipsoid, 15-26 × 13-18 mm, glabrous or sparsely puberulent, scar diam. 8-13 mm, scar often orangish.

biennial;

cup deeply and narrowly cup-shaped or U-shaped, 9-19 mm high × 7-18 mm wide, covering 1/3-1/2(-2/3) nut, outer surface glabrous to sparsely puberulent, inner surface glabrous or pubescent on innermost 1/3, occasionally uniformly pubescent, scales acute, tips loose;

nut narrowly conic or ovoid to narrowly oblong, 21-44 × 8-14 mm, glabrous, scar diam. 2.5-7.5 mm.

Terminal

buds gray to gray-brown, ovoid, 3-7 mm, glabrous or with scales somewhat ciliate at apex.

buds light chestnut brown to dark reddish brown, ovoid to conic, 3-9 mm, glabrous or with tuft of minute hairs at apex.

= 24.

Quercus texana

Quercus wislizenii

Phenology

Flowering spring.

Flowering late spring.

Habitat

Flood plains and bottomlands

Valleys, slopes, and sand chaparral

Elevation

0-200 m (0-700 ft)

300-1900 m (1000-6200 ft)

Distribution

AL; AR; IL; KY; LA; MO; MS; TN; TX

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

Discussion

For many years the name Quercus texana was erroneously used for Q. buckleyi (L. J. Dorr and K. C. Nixon 1985). A few authors have also used the name for Q. gravesii.

Quercus nuttallii E. J. Palmer var. cachensis E. J. Palmer was described as a small-fruited form (nuts 16-18 × 12-16 mm) from specimens collected in east-central Arkansas (E. J. Palmer 1937). Noting the similarity between Q. nuttallii var. cachensis and Q. palustris, Palmer discounted the possibility of the former being of hybrid origin because (1) he had not observed Q. palustris in the type locality, and (2) the leaves and buds of the former were essentially the same as in Q. nuttallii var. nuttallii.

C. H. Muller (1942), on the other hand, argued that Quercus nuttallii was nothing more than a form [forma nuttallii (E. J. Palmer) C. H. Muller] of Q. palustris. This is a puzzling conclusion because it was based largely on the premise that Q. nuttallii occurred "...with the parent species throughout a large part of the latter's southern range (Mississippi to eastern Texas and southeastern Missouri)." The range of Q. palustris does not extend into Mississippi or eastern Texas, although its range does overlap that of Q. texana in eastern Arkansas and southeastern Missouri. E. J. Palmer (1948) and D. M. Hunt (1989) have suggested hybridization with Q. shumardii and Q. nigra, respectively. See L. J. Dorr and K. C. Nixon (1985) for an explanation of the nomenclatural confusion regarding this taxon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Shrubs with oval leaves 25-38 mm and margins entire or deeply lobed-dentate may be treated as Quercus wislizenii var. frutescens. J. M. Tucker (1993) treated Q. parvula as a distinct species, distinguished from Q. wislizenii by its larger leaves (30-90 versus 20-50 mm), by the dull, olive-green, abaxial leaf surface (versus shiny, yellow-green), and by nuts that are abruptly tapered proximal to the middle (versus gradually tapered).

Tucker recognized two varieties of Quercus parvula: Q. parvula var. parvula is a shrub of 1-3 m and Q. parvula var. shrevei is a tree less than 17 m. S. K. Langer (1993) recognized a third variety, Q. parvula var. tamalpaisensis,based on several small populations on or near Mount Tamalpais, differentiated primarily by having larger leaves (50-160 × 20-60 mm) with attenuate-dentate margins.

Quercus wislizenii reportedly hybridizes with Q. agrifolia and Q. kelloggii (W. B. Brophy and D. R. Parnell 1974).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Lobatae

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana

Synonyms

Q. nuttallii, Q. rubra var. texana, Q. shumardii var. texana

Q. parvula, Q. parvula var. shrevei, Q. parvula var. tamalpaisensis, Q. wislizenii var. frutescens

Name authority

Buckley: Proc. Acad. Nat. Sci. Philadelphia 12: 444. (1860)

A. de Candolle: in A. P. de Candolle and A. L. P. de Candolle, Prodr. 16(2): 67. (1864)

Web links

Local Web sites: IL Wildflowers, LA Plants, MO Plants
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Quercus texana

Quercus wislizenii

Quercus texana

Quercus wislizenii