|
runner oak, running oak
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pungent oak, sandpaper oak
|
| Shrubs, deciduous or tardily deciduous, to 1 m. Bark gray to dark brown. |
Shrubs or moderate-sized trees, evergreen or subevergreen. |
|
light brown, papery. |
gray-brown to reddish brown, 1-2 mm diam., sparsely to uniformly pubescent. |
gray, 1-2 mm diam., short velvety-tomentose, glabrate with age. |
|
dark red-brown, ca. 2 mm, sparsely pubescent. |
blade oblong to narrowly obovate, 25-100 × 10-33 mm, base acute to rounded, margins entire, revolute, with 1 apical awn, apex acute or obtuse to rounded; surfaces abaxially uniformly gray-brown pubescent, rarely glabrate, adaxially somewhat convex, rugose, glabrous or with scattered hairs along midrib. |
blade elliptic to oblong, 10-40(-90) × 10-20(-40) mm, rather thick, leathery, stiff, base rounded or minutely cordate, very rarely cuneate, margins regularly undulate-crisped, not revolute, coarsely toothed or incised with acute teeth or spinose lobes, secondary veins 5-8(-14) on each side, usually branched before passing into teeth, apex acute or obtuse, rarely rounded, spine-tipped; surfaces abaxially canescent, usually densely stellate-pubescent, and mixed with stiff, harsh, stellate hairs, often sandpapery to touch, rarely glabrate, adaxially yellowish green, glossy, usually rough and sandpapery because of minute, persistent hair bases, rarely glabrate. |
annual; cup deeply saucer-shaped to turbinate, 5-12 mm high × 10-15 mm wide, covering 1/3-1/2(-2/3) nut, outer surface pubescent, inner surface densely pubescent, scales rarely involute, often tuberculate, tips tightly appressed, acute; nut globose to ovoid or broadly oblong, 9.5-15 × 9-12 mm, glabrate, scar diam. 5-8 mm. |
subsessile or on peduncle to 3 mm; cup shallowly to deeply cup-shaped or turbinate, to 8 mm deep × 13 mm wide, covering ca. 1/4 nut, margin thin, scales reddish brown, moderately tuberculate or keeled, densely gray-tomentose; nut light brown, broadly ovoid to subcylindric, to 10 × 10 mm, apex rounded to subacute, glabrous. |
|
distinct. |
buds brown to red-brown, ovoid, 2.5-4.5 mm, glabrous or with ciliate scale margins. |
|
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| Flowering spring. |
Flowering spring. |
| Dry sandy soils of savannahs, low ridges and oak-pine scrub, occasionally at margins of poorly drained sites |
On dry limestone or igneous slopes, usually in oak, pinyon, and juniper woodlands, chaparral, and sometimes descending into desert vegetation |
| 0-100 m (0-300 ft) |
800-2000 m (2600-6600 ft) |
|
AL; FL; GA; MS; NC; SC
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AZ; NM; TX; Mexico (Chihuahua and Coahuila)
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Although no hybrid combinations have been formally proposed, D. M. Hunt (1989) has reported evidence of hybridization with Quercus hemisphaerica, Q. incana, Q. myrtifolia, and Q. phellos. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Numerous populations appear to be hybrid swarms between Quercus pungens and Q. vaseyana, which is sometimes treated as a variety of Q. pungens. No other evidence for a close relationship exists for these two species, and such a treatment risks erecting a polyphyletic assemblage. To the west and south within the range of Q. pungens no indication of introgression exists, and the two species are strikingly different and easily separable. I interpret the contact as secondary. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 3. |
FNA vol. 3. |
| Fagaceae > Quercus > sect. Lobatae |
Fagaceae > Quercus > sect. Quercus |
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii |
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii |
|
Q. undulata var. pungens |
| Walter: Fl. Carol., 234. (1788) |
Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171. 185: not Q. pungens Gandoger 1890 |
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