FNA: Quercus pumila vs. Quercus montana

	Quercus pumila	Quercus montana
	runner oak, running oak	chestnut oak, mountain chestnut oak, rock chestnut oak
Habit	Shrubs, deciduous or tardily deciduous, to 1 m. Bark gray to dark brown.	Trees, deciduous, to 30 m. Bark dark gray or brown, hard, with deep V-shaped furrows.
Twigs	gray-brown to reddish brown, 1-2 mm diam., sparsely to uniformly pubescent.	light brown, 2-3(-4) mm diam., glabrous.
Buds		light brown, ovoid, (3-)4-6 mm, occasionally apex acute, glabrous.
Leaves	blade oblong to narrowly obovate, 25-100 × 10-33 mm, base acute to rounded, margins entire, revolute, with 1 apical awn, apex acute or obtuse to rounded; surfaces abaxially uniformly gray-brown pubescent, rarely glabrate, adaxially somewhat convex, rugose, glabrous or with scattered hairs along midrib.	blade obovate to narrowly elliptic or narrowly obovate, (100-)120-200(-220) × 60-100(-120) mm, base subacute or rounded-acuminate, often unequal, margins regularly toothed, teeth rounded or rarely somewhat acute, secondary veins ± parallel, straight or moderately curved, 10-14(-16) on each side, apex broadly acuminate; surfaces abaxially light green, appearing glabrous but with scattered minute, asymmetric, appressed-stellate hairs and usually visible, larger, simple or fascicled erect hairs along veins, adaxially dark green, glossy, glabrous or with minute, scattered, simple hairs.
Acorns	annual; cup deeply saucer-shaped to turbinate, 5-12 mm high × 10-15 mm wide, covering 1/3-1/2(-2/3) nut, outer surface pubescent, inner surface densely pubescent, scales rarely involute, often tuberculate, tips tightly appressed, acute; nut globose to ovoid or broadly oblong, 9.5-15 × 9-12 mm, glabrate, scar diam. 5-8 mm.	1-3, subsessile or on peduncle 8-20(-25) mm; cup shallowly cup-shaped to hemispheric or deeply goblet-shaped, rim thin, often flared and undulate, helmetlike, 9-15 mm deep × 18-25 mm wide, scales often in concentric or transverse rows, laterally connate, gray, broadly ovate, tips reddish, glabrous; nut light brown, ovoid-ellipsoid, 15-30 × 10-20(-25) mm, glabrous.
Cotyledons		distinct.
Terminal	buds brown to red-brown, ovoid, 2.5-4.5 mm, glabrous or with ciliate scale margins.
2n		= 24.

	Quercus pumila	Quercus montana
Phenology	Flowering spring.	Flowering mid-late spring.
Habitat	Dry sandy soils of savannahs, low ridges and oak-pine scrub, occasionally at margins of poorly drained sites	Rocky upland forest, dry ridges, mixed deciduous forests on shallow soils
Elevation	0-100 m (0-300 ft)	0-1400 m (0-4600 ft)
Distribution	AL; FL; GA; MS; NC; SC [WildflowerSearch map]	AL; CT; DE; GA; IL; IN; KY; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV [WildflowerSearch map] [BONAP county map]
Discussion	Although no hybrid combinations have been formally proposed, D. M. Hunt (1989) has reported evidence of hybridization with Quercus hemisphaerica, Q. incana, Q. myrtifolia, and Q. phellos. (Discussion copyrighted by Flora of North America; reprinted with permission.)	The name Quercus prinus Linnaeus is often applied to this species, particularly in the forestry literature, and in many regional floras. In a number of works, however, Q. prinus has been applied to the species here treated as Q. michauxii. Following the recommendations of J. W. Hardin (1979), because of the persistent confusion in the application of the name Q. prinus and uncertainty regarding the identity of the Linnean type materials, the names Q. montana and Q. michauxii should be used for the two species that have been variously called Q. prinus. Quercus prinus under this interpretation is a name of uncertain position. The four species of the chestnut oak group in eastern North America (Quercus montana, Q. michauxii, Q. muhlenbergii, and Q. prinoides) are somewhat difficult to distinguish unless careful attention is paid to features of leaf vestiture and fruit and cup morphology. Attempts to identify these species mostly or solely on basis of leaf shape and dentition (as in many other oak species complexes) have resulted in a plethora of misidentified material in herbaria and erroneous reports in the literature. The closely appressed, asymmetric trichomes on the abaxial surface of the mature leaf, in combination with longer simple hairs along the midvein, are unique to Q. montana among North American species of Quercus. Immature leaves and densely shaded leaves sometimes exhibit a more erect trichome that could be confused with the longer, felty hairs of Q. michauxii, so it is important to evaluate mature sun leaves when possible. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Lobatae	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Name authority	Walter: Fl. Carol., 234. (1788)	Willdenow: Sp. Pl. 4(1): 440. (1805)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: Go Botany, IL Wildflowers, MD Biodiversity, MI Flora WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus montana

runner oak, running oak

chestnut oak, mountain chestnut oak, rock chestnut oak

Habit

Shrubs, deciduous or tardily deciduous, to 1 m. Bark gray to dark brown.

Trees, deciduous, to 30 m. Bark dark gray or brown, hard, with deep V-shaped furrows.

Twigs

gray-brown to reddish brown, 1-2 mm diam., sparsely to uniformly pubescent.

light brown, 2-3(-4) mm diam., glabrous.

Buds

light brown, ovoid, (3-)4-6 mm, occasionally apex acute, glabrous.

Leaves

blade oblong to narrowly obovate, 25-100 × 10-33 mm, base acute to rounded, margins entire, revolute, with 1 apical awn, apex acute or obtuse to rounded;

surfaces abaxially uniformly gray-brown pubescent, rarely glabrate, adaxially somewhat convex, rugose, glabrous or with scattered hairs along midrib.

blade obovate to narrowly elliptic or narrowly obovate, (100-)120-200(-220) × 60-100(-120) mm, base subacute or rounded-acuminate, often unequal, margins regularly toothed, teeth rounded or rarely somewhat acute, secondary veins ± parallel, straight or moderately curved, 10-14(-16) on each side, apex broadly acuminate;

surfaces abaxially light green, appearing glabrous but with scattered minute, asymmetric, appressed-stellate hairs and usually visible, larger, simple or fascicled erect hairs along veins, adaxially dark green, glossy, glabrous or with minute, scattered, simple hairs.

Acorns

annual;

cup deeply saucer-shaped to turbinate, 5-12 mm high × 10-15 mm wide, covering 1/3-1/2(-2/3) nut, outer surface pubescent, inner surface densely pubescent, scales rarely involute, often tuberculate, tips tightly appressed, acute;

nut globose to ovoid or broadly oblong, 9.5-15 × 9-12 mm, glabrate, scar diam. 5-8 mm.

1-3, subsessile or on peduncle 8-20(-25) mm;

cup shallowly cup-shaped to hemispheric or deeply goblet-shaped, rim thin, often flared and undulate, helmetlike, 9-15 mm deep × 18-25 mm wide, scales often in concentric or transverse rows, laterally connate, gray, broadly ovate, tips reddish, glabrous;

nut light brown, ovoid-ellipsoid, 15-30 × 10-20(-25) mm, glabrous.

Cotyledons

distinct.

Terminal

buds brown to red-brown, ovoid, 2.5-4.5 mm, glabrous or with ciliate scale margins.

= 24.

Quercus pumila

Quercus montana

Phenology

Flowering spring.

Flowering mid-late spring.

Habitat

Dry sandy soils of savannahs, low ridges and oak-pine scrub, occasionally at margins of poorly drained sites

Rocky upland forest, dry ridges, mixed deciduous forests on shallow soils

Elevation

0-100 m (0-300 ft)

0-1400 m (0-4600 ft)

Distribution

AL; FL; GA; MS; NC; SC

[WildflowerSearch map]

AL; CT; DE; GA; IL; IN; KY; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV

[WildflowerSearch map]

[BONAP county map]

Discussion

Although no hybrid combinations have been formally proposed, D. M. Hunt (1989) has reported evidence of hybridization with Quercus hemisphaerica, Q. incana, Q. myrtifolia, and Q. phellos.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Quercus prinus Linnaeus is often applied to this species, particularly in the forestry literature, and in many regional floras. In a number of works, however, Q. prinus has been applied to the species here treated as Q. michauxii. Following the recommendations of J. W. Hardin (1979), because of the persistent confusion in the application of the name Q. prinus and uncertainty regarding the identity of the Linnean type materials, the names Q. montana and Q. michauxii should be used for the two species that have been variously called Q. prinus. Quercus prinus under this interpretation is a name of uncertain position.

The four species of the chestnut oak group in eastern North America (Quercus montana, Q. michauxii, Q. muhlenbergii, and Q. prinoides) are somewhat difficult to distinguish unless careful attention is paid to features of leaf vestiture and fruit and cup morphology. Attempts to identify these species mostly or solely on basis of leaf shape and dentition (as in many other oak species complexes) have resulted in a plethora of misidentified material in herbaria and erroneous reports in the literature. The closely appressed, asymmetric trichomes on the abaxial surface of the mature leaf, in combination with longer simple hairs along the midvein, are unique to Q. montana among North American species of Quercus. Immature leaves and densely shaded leaves sometimes exhibit a more erect trichome that could be confused with the longer, felty hairs of Q. michauxii, so it is important to evaluate mature sun leaves when possible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Lobatae

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Name authority

Walter: Fl. Carol., 234. (1788)

Willdenow: Sp. Pl. 4(1): 440. (1805)

Web links

Local Web sites: Go Botany, IL Wildflowers, MD Biodiversity, MI Flora
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Quercus pumila

Quercus montana

Quercus pumila

Quercus montana