Quercus palmeri |
Quercus velutina |
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Palmer oak, Palmer's oak |
black oak |
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Habit | Trees, deciduous, to 25 m. Bark dark brown to black, deeply furrowed, ridges often broken into irregular blocks, inner bark yellow or orange. | |
Twigs | dark reddish brown, (1.5-)2.5-4.5(-5) mm diam., glabrous or sparsely pubescent. |
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Leaves | blade suborbiculate, elliptic to round-ovate, 20-30(-50) × 20-40 mm, crisped, leathery and brittle, base obtuse to strongly subcordate, secondary veins 5-8(-12) pairs, each terminating in spine, basal pairs recurving, others branching at 45° angles, raised abaxially, margins spinose-dentate to occasionally entire, with highly thickened cell walls, spines cartilaginous, (1-)1.5-2 mm, apex broadly rounded or subacute, spinose; surfaces abaxially glaucous with waxy layer, often obscured by golden brown glandular hairs, adaxially grayish dark green, scurfy with fasciculate erect and twisting hairs. |
blade ovate to obovate, (80-)100-300 × 80-150 mm, base obtuse to truncate, inequilateral, margins with 5-9 lobes and 15-50 awns, lobes oblong or distally expanded, separated by deep sinuses, apex acute to obtuse; surfaces abaxially pale green, glabrous except for small axillary tufts of tomentum or with scattered pubescence, especially along veins, adaxially glossy, dark green, glabrous, secondary veins raised on both surfaces. |
Acorns | solitary or rarely paired; cup turbinate to saucer-shaped, margins involute, often irregular, 7-10 mm deep × 10-25(-35) mm wide, scales appressed, embedded, often appearing laterally connate into concentric rings with only tip of scale visible, tuberculate, densely golden-tomentose throughout; nut oblong to fusiform, 20-30 × 10-15 mm, apex acute. |
biennial; cup cup-shaped or turbinate, 7-14 mm high × 12-22 mm wide, covering 1/2 nut, cup margins not involute, outer surface puberulent, inner surface pubescent, scale tips loose, especially at margin of cup, acute to acuminate; nut subglobose to ovoid, 10-20 × 10-18 mm, glabrate, scar diam. 5.5-12 mm. |
Small | trees and shrubs, to 2-3 m. Twigs rigid, divaricately branched at 65-90° angles, reddish brown, 1.5-3 mm diam., pubescent, sparsely so in 2d year. |
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Terminal | buds ovoid, 1-1.5 mm, apex rounded, glabrous. |
buds ovoid or ellipsoid to subconic, 6-12 mm, noticeably 5-angled in cross section, tawny- or gray-pubescent. |
n | = 12 ± 1; 2n = 24. |
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Quercus palmeri |
Quercus velutina |
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Phenology | Flowering in spring. | Flowering spring. |
Habitat | Disjunct in canyons, mountain washes, dry thickets, and margins of chapparal communities | Commonly on dry slopes and upland areas, occasionally on sandy lowlands (especially in north) and poorly drained uplands and terraces |
Elevation | 700-1800 m (2300-5900 ft) | 0-1500 m (0-4900 ft) |
Distribution |
AZ; CA; Mexico (n Baja California)
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON
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Discussion | Populations of Quercus palmeri are often small and may exist as single clones. The disjunct populations of California and Baja California are consistent morphologically. In Arizona populations, individuals tend to have flatter leaves bearing fewer teeth; this distinction is not constant, however. Morphologically aberrant populations identified as Q. palmeri in eastern Arizona (Chiracahua, Huachuca, and Santa Catalina mountains) and southwestern New Mexico are most likely the result of introgression from Q. palmeri to Q. chrysolepis (J. M. Tucker and H. S. Haskell 1960). Those populations tend to be intermediate in overall morphology, but all lack the diagnostic trichomes and biochemical markers of Q. palmeri; they are best classified as Q. chrysolepis affinity Q. palmeri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The bark of this species (quercitron) is rich in tannins and was once an important source of these chemicals used for tanning leather. (The yellow dye obtained from the bark is also called quercitron.) Native Americans used Quercus velutina medicinally for indigestion, chronic dysentery, mouth sores, chills and fevers, chapped skin, hoarseness, milky urine, lung trouble, sore eyes, and as a tonic, an antiseptic, and an emetic (D. E. Moerman 1986). Quercus velutina reportedly hybridizes with Q. coccinea, Q. ellipsoidalis (= Q. ×paleolithicola Trelease), Q. falcata [= Q. ×willdenowiana (Dippel) Zabel] (= Q. ×pinetorum Moldenke)], Q. ilicifolia (= Q. ×rehderi Trelease), Q. imbricaria (= Q. ×leana Nuttall), Q. incana, Q. laevis, and Q. laurifolia (= Q. ×cocksii Sargent, although E. J. Palmer [1948] challenged the validity of this claim), Q. marilandica, Q. nigra, Q. palustris (= Q. ×vaga E. J. Palmer & Steyermark), Q. phellos (= Q. ×filialis Little), Q. rubra, Q. shumardii, and possibly Q. arkansana (D. M. Hunt 1989). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Protobalanus | Fagaceae > Quercus > sect. Lobatae |
Sibling taxa | ||
Synonyms | Q. dunnii | Q. tinctoria |
Name authority | Engelmann: in S. Watson, Bot. California 2: 97. (1880) | Lamarck: in J. Lamarck et al., Encycl. 1: 721. (1785) |
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