FNA: Quercus palmeri vs. Quercus depressipes

	Quercus palmeri	Quercus depressipes
	Palmer oak, Palmer's oak	Davis Mountain oak, depressed oak
Habit		Shrubs, evergreen or subevergreen, low, to 1 m, often forming dense thickets, rhizomatous.
Bark		gray, scaly.
Twigs		tan-brown, becoming reddish gray, 1-1.5 mm diam., glabrate or hairy.
Buds		tan or brown, subglobose, 1-1.5 mm, glabrate or scales inconspicuously ciliate.
Leaves	blade suborbiculate, elliptic to round-ovate, 20-30(-50) × 20-40 mm, crisped, leathery and brittle, base obtuse to strongly subcordate, secondary veins 5-8(-12) pairs, each terminating in spine, basal pairs recurving, others branching at 45° angles, raised abaxially, margins spinose-dentate to occasionally entire, with highly thickened cell walls, spines cartilaginous, (1-)1.5-2 mm, apex broadly rounded or subacute, spinose; surfaces abaxially glaucous with waxy layer, often obscured by golden brown glandular hairs, adaxially grayish dark green, scurfy with fasciculate erect and twisting hairs.	blade oblong to elliptic, 10-25(-60) × 8-25 mm, thick, leathery, base moderately to deeply cordate, petiole strongly depressed in basal sinus, margins inconspicuously toothed in distal 1/2, rarely entire, sometimes sublobate, somewhat revolute, secondary veins 5 or 6 on each side with few intermediates, branching, apex broadly rounded to subacute; surfaces abaxially dull gray-green or glaucous, completely glabrous or with a few stellate hairs on midrib, adaxially similar to abaxial surface, secondary veins somewhat raised on both surfaces.
Acorns	solitary or rarely paired; cup turbinate to saucer-shaped, margins involute, often irregular, 7-10 mm deep × 10-25(-35) mm wide, scales appressed, embedded, often appearing laterally connate into concentric rings with only tip of scale visible, tuberculate, densely golden-tomentose throughout; nut oblong to fusiform, 20-30 × 10-15 mm, apex acute.	paired on peduncle 7-15 mm; cup 4-7 mm deep × 8-13 mm wide, goblet-shaped, enclosing 1/4-1/2 nut, base somewhat constricted or rounded, scales moderately tuberculate, proximally densely gray-tomentose, tips rather closely appressed, reddish brown, abaxially glabrous, ciliate; nut tan-brown, elliptic to ovoid or globose, to 10-15 × 10-11 mm, apex rounded, glabrous.
Cotyledons		connate.
Small	trees and shrubs, to 2-3 m. Twigs rigid, divaricately branched at 65-90° angles, reddish brown, 1.5-3 mm diam., pubescent, sparsely so in 2d year.
Terminal	buds ovoid, 1-1.5 mm, apex rounded, glabrous.

	Quercus palmeri	Quercus depressipes
Phenology	Flowering in spring.	Flowering spring.
Habitat	Disjunct in canyons, mountain washes, dry thickets, and margins of chapparal communities	Grassland and open wooded slopes
Elevation	700-1800 m (2300-5900 ft)	2100-2600 m (6900-8500 ft)
Distribution	AZ; CA; Mexico (n Baja California) [WildflowerSearch map] [BONAP county map]	TX; Mexico (Chihuahua, Durango, and Zacatecas) [BONAP county map]
Discussion	Populations of Quercus palmeri are often small and may exist as single clones. The disjunct populations of California and Baja California are consistent morphologically. In Arizona populations, individuals tend to have flatter leaves bearing fewer teeth; this distinction is not constant, however. Morphologically aberrant populations identified as Q. palmeri in eastern Arizona (Chiracahua, Huachuca, and Santa Catalina mountains) and southwestern New Mexico are most likely the result of introgression from Q. palmeri to Q. chrysolepis (J. M. Tucker and H. S. Haskell 1960). Those populations tend to be intermediate in overall morphology, but all lack the diagnostic trichomes and biochemical markers of Q. palmeri; they are best classified as Q. chrysolepis affinity Q. palmeri. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Quercus depressipes enters the United States in only one population on the highest portion of Mt. Livermore in trans-Pecos Texas; it has a wider distribution in the dry altiplano of northern Mexico. Its most distinctive characteristics are the combination of dwarf clonal habit, small glaucous leaves without spinose teeth, and connate cotyledons. In northern Mexico, it hybridizes locally with Q. rugosa. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Protobalanus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. dunnii	Q. bocoynensis
Name authority	Engelmann: in S. Watson, Bot. California 2: 97. (1880)	Trelease: Mem. Natl. Acad. Sci. 20: 90. (1924)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus depressipes

Palmer oak, Palmer's oak

Davis Mountain oak, depressed oak

Habit

Shrubs, evergreen or subevergreen, low, to 1 m, often forming dense thickets, rhizomatous.

Bark

gray, scaly.

Twigs

tan-brown, becoming reddish gray, 1-1.5 mm diam., glabrate or hairy.

Buds

tan or brown, subglobose, 1-1.5 mm, glabrate or scales inconspicuously ciliate.

Leaves

blade suborbiculate, elliptic to round-ovate, 20-30(-50) × 20-40 mm, crisped, leathery and brittle, base obtuse to strongly subcordate, secondary veins 5-8(-12) pairs, each terminating in spine, basal pairs recurving, others branching at 45° angles, raised abaxially, margins spinose-dentate to occasionally entire, with highly thickened cell walls, spines cartilaginous, (1-)1.5-2 mm, apex broadly rounded or subacute, spinose;

surfaces abaxially glaucous with waxy layer, often obscured by golden brown glandular hairs, adaxially grayish dark green, scurfy with fasciculate erect and twisting hairs.

blade oblong to elliptic, 10-25(-60) × 8-25 mm, thick, leathery, base moderately to deeply cordate, petiole strongly depressed in basal sinus, margins inconspicuously toothed in distal 1/2, rarely entire, sometimes sublobate, somewhat revolute, secondary veins 5 or 6 on each side with few intermediates, branching, apex broadly rounded to subacute;

surfaces abaxially dull gray-green or glaucous, completely glabrous or with a few stellate hairs on midrib, adaxially similar to abaxial surface, secondary veins somewhat raised on both surfaces.

Acorns

solitary or rarely paired;

cup turbinate to saucer-shaped, margins involute, often irregular, 7-10 mm deep × 10-25(-35) mm wide, scales appressed, embedded, often appearing laterally connate into concentric rings with only tip of scale visible, tuberculate, densely golden-tomentose throughout;

nut oblong to fusiform, 20-30 × 10-15 mm, apex acute.

paired on peduncle 7-15 mm;

cup 4-7 mm deep × 8-13 mm wide, goblet-shaped, enclosing 1/4-1/2 nut, base somewhat constricted or rounded, scales moderately tuberculate, proximally densely gray-tomentose, tips rather closely appressed, reddish brown, abaxially glabrous, ciliate;

nut tan-brown, elliptic to ovoid or globose, to 10-15 × 10-11 mm, apex rounded, glabrous.

Cotyledons

connate.

Small

trees and shrubs, to 2-3 m. Twigs rigid, divaricately branched at 65-90° angles, reddish brown, 1.5-3 mm diam., pubescent, sparsely so in 2d year.

Terminal

buds ovoid, 1-1.5 mm, apex rounded, glabrous.

Quercus palmeri

Quercus depressipes

Phenology

Flowering in spring.

Flowering spring.

Habitat

Disjunct in canyons, mountain washes, dry thickets, and margins of chapparal communities

Grassland and open wooded slopes

Elevation

700-1800 m (2300-5900 ft)

2100-2600 m (6900-8500 ft)

Distribution

AZ; CA; Mexico (n Baja California)

[WildflowerSearch map]

[BONAP county map]

TX; Mexico (Chihuahua, Durango, and Zacatecas)

[BONAP county map]

Discussion

Populations of Quercus palmeri are often small and may exist as single clones. The disjunct populations of California and Baja California are consistent morphologically. In Arizona populations, individuals tend to have flatter leaves bearing fewer teeth; this distinction is not constant, however. Morphologically aberrant populations identified as Q. palmeri in eastern Arizona (Chiracahua, Huachuca, and Santa Catalina mountains) and southwestern New Mexico are most likely the result of introgression from Q. palmeri to Q. chrysolepis (J. M. Tucker and H. S. Haskell 1960). Those populations tend to be intermediate in overall morphology, but all lack the diagnostic trichomes and biochemical markers of Q. palmeri; they are best classified as Q. chrysolepis affinity Q. palmeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus depressipes enters the United States in only one population on the highest portion of Mt. Livermore in trans-Pecos Texas; it has a wider distribution in the dry altiplano of northern Mexico. Its most distinctive characteristics are the combination of dwarf clonal habit, small glaucous leaves without spinose teeth, and connate cotyledons. In northern Mexico, it hybridizes locally with Q. rugosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Protobalanus

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. dunnii

Q. bocoynensis

Name authority

Engelmann: in S. Watson, Bot. California 2: 97. (1880)

Trelease: Mem. Natl. Acad. Sci. 20: 90. (1924)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus palmeri

Quercus depressipes

Quercus palmeri

Quercus depressipes