Flora of North America species comparison

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Quercus pagoda

Quercus havardii

cherrybark oak, Texas oak

Havard oak, Havard's oak, Havard's shinnery oak, sand shinnery oak, sand shinoak, shinnery oak
Habit Trees, deciduous, to 40 m. Bark nearly black with narrow and noticeably flaky ridges, often resembling that of wild black cherry, inner bark orange. Shrubs, deciduous, low, forming clones 0.3-1.5 × 10 m, rhizomatous.
Bark

light gray, scaly-papery.
Twigs

yellowish brown, 2-3.5 mm diam., pubescent.

brown or grayish, 1-2.5 mm diam., glabrous or densely short grayish or yellowish tomentulose, glabrate in age.
Buds

dark red-brown, subglobose, ca. 2 mm, sparsely pubescent.
Leaves

blade ovate to elliptic or obovate, 90-300 × 60-160 mm, base cuneate to rounded or truncate, margins with 5-11 lobes and 10-25 awns, lobes oblong, rarely falcate, terminal lobe rarely exceeding lateral lobes in length, apex acute;

surfaces abaxially pale, tomentose, adaxially glossy, glabrous, secondary veins raised on both surfaces.

blade green, often turning brownish with age, polymorphic, oblong or elliptic or sometimes lanceolate to oblanceolate or ovate to obovate, (30-)50-100 × (10-)20-50 mm, rather thick and hard, base rounded to cuneate, margins flat to revolute, at least some undulate, 2-3 rounded teeth on each side, secondary veins 5-8 on each side, much branched, apex broadly rounded, rarely acute;

surfaces abaxially densely grayish or yellowish tomentulose or stellate-pubescent, sometimes only sparsely pubescent, secondary veins quite prominent, adaxially lustrous, very sparsely stellate-pubescent or glabrate, secondary veins very slightly if at all raised.
Acorns

biennial;

cup saucer-shaped to cup-shaped, 3-7 mm high × 10-18 mm wide, covering 1/3-1/2 nut, outer surface puberulent, inner surface pubescent, scale tips tightly appressed, acute;

nut subglobose, 9-15 × 8-16 mm, often striate, puberulent, scar diam. 5-9 mm.

solitary or paired, subsessile or on peduncle to 10(-18) mm;

cup from deeply cup-shaped to goblet-shaped, 10-12 mm deep × 15-25 mm wide, enclosing 1/3-1/2 nut, base rounded or slightly constricted, margin very thin and smooth, scales reddish brown, triangular-ovate to long-acute, proximally moderately to markedly tuberculate, pubescent, often canescent, tips loosely appressed;

nut brown, ovoid, 12-25 × 14-18 mm.
Cotyledons

distinct.
Terminal

buds light reddish brown, ovoid, 4-9 mm, strongly 5-angled in cross section, puberulent throughout.

Quercus pagoda

Quercus havardii
Phenology Flowering spring. Flowering spring.
Habitat Poorly drained bottoms and mesic slopes Deep, shifting or stabilized sand dunes, off deep sands in putative hybrid populations
Elevation 0-300 m (0-1000 ft) 500-1500 m (1600-4900 ft)
Distribution
from FNA
AL; AR; FL; GA; IL; IN; KY; LA; MO; MS; NC; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Quercus pagoda is often treated as a variety of Q. falcata; it is quite distinctive, however, both morphologically and ecologically (S. A. Ware 1967; R. J. Jensen 1989).

This species reportedly hybridizes with Q. falcata and Q. phellos (D. M. Hunt 1989).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Individual clones emerging to heights of 2-3 m from thickets occur sporadically across the Texas range of Quercus havardii and express some characteristics of Q. stellata, such as more deeply lobed leaves and smaller acorns. Such putative hybrids increase in frequency in the eastern part of the range of the species.

Material of Quercus havardii from the Navajo Basin of Utah and adjacent Arizona has been treated as Q. havardii var. tuckeri Welsh. Welsh followed J. M. Tucker (1970) and interpreted these intermediate populations as putative hybrids between Q. havardii and both Q. turbinella and Q. gambelii. Giving varietal rank, instead of nothospecies status, to such populations seems arbitrary, and it certainly is inconsistent with their putative hybrid origins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Lobatae Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. falcata var. leucophylla, Q. falcata var. pagodifolia, Q. leucophylla, Q. pagodifolia
Name authority Rafinesque: Alsogr. Amer., 23. (1838) Rydberg: Bull. New York Bot. Gard. 2: 213. (1901)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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