Quercus pacifica |
Quercus vacciniifolia |
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Channel Island scrub oak, island scrub oak, Pacific oak |
huckleberry oak |
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Habit | Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller. | Shrubs, low spreading to often prostrate, to 1.5 m. Twigs branching at 45° angles or less, reddish brown, 1-1.5 mm diam., flexible, glabrous to sparsely pubesent. |
Bark | scaly on older branches and trunk. |
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Twigs | brownish or reddish, minutely puberulent, becoming glabrate and gray with age. |
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Buds | light or chestnut brown, ovate or globose, 2-3 × 1-2 mm. |
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Leaves | blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip; surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs. |
blade oblong-ovate, 10-35 × 7-15 mm, flat, thin, leathery, base slightly rounded to acute, secondary veins inconspicuous, 6-8 pairs, branching at 45-60° angles, with weakly thickened cell walls, margins entire or indistinctly and irregularly mucronately toothed, apex acute or rarely obtuse; surfaces abaxially whitish green with waxy layer, glabrous or slightly pubescent with stellate hairs, adaxially dull gray-green, glabrous or sparsely pubescent with stellate hairs. |
Acorns | paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms; cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed; nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate. |
solitary or rarely paired; cup shallowly saucer-shaped to slightly turbinate, 3-4 mm deep × 10-15 mm wide, scales appressed, slightly embedded, moderately silvery brown-pubescent; nut ovoid, 8-17 × 5-10 mm, apex acute; nut scar to 3 mm diam. |
Cotyledons | distinct. |
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Terminal | buds conic, 2.5 mm, scales brown with ciliate margins. |
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Quercus pacifica |
Quercus vacciniifolia |
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Phenology | Flowering spring. | Flowering in early summer. |
Habitat | Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands | Dry ridges, steep slopes, and rocky areas from montane coniferous zone to near treeline |
Elevation | 0-300 m (0-1000 ft) | 900-2800 m (3000-9200 ft) |
Distribution |
CA
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CA; NV; OR
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Discussion | Of conservation concern. Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands. Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Typical high-elevation populations in the Sierra Nevada of California can be distinguished from all shrubby forms of Quercus chrysolepis by the absence of glandular trichomes and by thin cups with small nut-attachment scars. At lower elevations in northern California and southwestern Oregon, secondary contact with Q. chrysolepis has resulted in the formation of hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Quercus | Fagaceae > Quercus > sect. Protobalanus |
Sibling taxa | ||
Synonyms | Q. dumosa var. polycarpa | |
Name authority | Nixon & C. H. Muller: Novon 4: 391. (1994) | Hittell: Resources Calif. 101. (1863) — (as vaccinifolia) |
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