FNA: Quercus pacifica vs. Quercus rugosa

	Quercus pacifica	Quercus rugosa
	Channel Island scrub oak, island scrub oak, Pacific oak	netleaf oak
Habit	Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller.	Shrubs or trees, evergreen, usually moderate-sized, rarely large.
Bark	scaly on older branches and trunk.	light or dark brown, scaly.
Twigs	brownish or reddish, minutely puberulent, becoming glabrate and gray with age.	brown, turning gray with age, 1-2 mm diam., tomentose to tomentulose, variously glabrate or persistently pubescent.
Buds	light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.	brown, ovoid, 2-4 mm, apex obtuse, sparsely pubescent or eventually glabrate.
Leaves	blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip; surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.	blade broadly obovate or panduriform to orbiculate or elliptic, rarely narrowly obovate, usually cupped, strongly concave proximally, sometimes planar, to 100 × 70 mm, stiff, leathery, base deeply or shallowly cordate, margins usually somewhat revolute, cartilaginously thickened, undulately crisped or flat with inconspicuous or coarse mucronate teeth near apex, secondary veins 8-10(-12) on each side, branched, apex broadly rounded, rarely subacute; surfaces abaxially dull, glaucous, or densely brownish tomentose, becoming nearly glabrate or pubescence persistent, especially about midribs, secondary veins very prominently raised, reticulate, adaxially dark green, lustrous, sparsely stellate-pubescent especially about base of midrib, secondary veins impressed.
Acorns	paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms; cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed; nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.	1-3 or more on slender axillary peduncle 30-60 mm; cup deeply cup-shaped to saucer-shaped, to 9 mm deep × 15 mm wide, enclosing to 1/2 nut, scales loosely appressed, characteristically somewhat spreading, brown, ovate, tuberculate-thickened or only slightly so, tomentose or obscurely tomentulose; nut light brown, ovoid to elliptic, to 20 × 15 mm, glabrous or minutely villous.
Cotyledons	distinct.	distinct, often reddish or purple.

	Quercus pacifica	Quercus rugosa
Phenology	Flowering spring.	Flowering early–late spring.
Habitat	Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands	Wooded slopes
Elevation	0-300 m (0-1000 ft)	2000-2500 m (6600-8200 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	AZ; NM; TX; Mexico; Central America (Guatemala) [WildflowerSearch map] [BONAP county map]
Discussion	Of conservation concern. Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands. Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Quercus rugosa occurs on wooded slopes at high elevations in trans-Pecos Texas, southern New Mexico, and Arizona, and throughout most of the mesic montane parts of Mexico, south to Guatemala. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. dumosa var. polycarpa	Q. ariifolia, Q. diversicolor, Q. durangensis, Q. reticulata, Q. rhodophlebia, Q. vellifera
Name authority	Nixon & C. H. Muller: Novon 4: 391. (1994)	Nee.: Anales Ci. Nat. 3: 275. (1801)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus rugosa

Channel Island scrub oak, island scrub oak, Pacific oak

netleaf oak

Habit

Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller.

Shrubs or trees, evergreen, usually moderate-sized, rarely large.

Bark

scaly on older branches and trunk.

light or dark brown, scaly.

Twigs

brownish or reddish, minutely puberulent, becoming glabrate and gray with age.

brown, turning gray with age, 1-2 mm diam., tomentose to tomentulose, variously glabrate or persistently pubescent.

Buds

light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.

brown, ovoid, 2-4 mm, apex obtuse, sparsely pubescent or eventually glabrate.

Leaves

blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip;

surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.

blade broadly obovate or panduriform to orbiculate or elliptic, rarely narrowly obovate, usually cupped, strongly concave proximally, sometimes planar, to 100 × 70 mm, stiff, leathery, base deeply or shallowly cordate, margins usually somewhat revolute, cartilaginously thickened, undulately crisped or flat with inconspicuous or coarse mucronate teeth near apex, secondary veins 8-10(-12) on each side, branched, apex broadly rounded, rarely subacute;

surfaces abaxially dull, glaucous, or densely brownish tomentose, becoming nearly glabrate or pubescence persistent, especially about midribs, secondary veins very prominently raised, reticulate, adaxially dark green, lustrous, sparsely stellate-pubescent especially about base of midrib, secondary veins impressed.

Acorns

paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms;

cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed;

nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.

1-3 or more on slender axillary peduncle 30-60 mm;

cup deeply cup-shaped to saucer-shaped, to 9 mm deep × 15 mm wide, enclosing to 1/2 nut, scales loosely appressed, characteristically somewhat spreading, brown, ovate, tuberculate-thickened or only slightly so, tomentose or obscurely tomentulose;

nut light brown, ovoid to elliptic, to 20 × 15 mm, glabrous or minutely villous.

Cotyledons

distinct.

distinct, often reddish or purple.

Quercus pacifica

Quercus rugosa

Phenology

Flowering spring.

Flowering early–late spring.

Habitat

Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands

Wooded slopes

Elevation

0-300 m (0-1000 ft)

2000-2500 m (6600-8200 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

AZ; NM; TX; Mexico; Central America (Guatemala)

[WildflowerSearch map]

[BONAP county map]

Discussion

Of conservation concern.

Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands.

Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus rugosa occurs on wooded slopes at high elevations in trans-Pecos Texas, southern New Mexico, and Arizona, and throughout most of the mesic montane parts of Mexico, south to Guatemala.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. dumosa var. polycarpa

Q. ariifolia, Q. diversicolor, Q. durangensis, Q. reticulata, Q. rhodophlebia, Q. vellifera

Name authority

Nixon & C. H. Muller: Novon 4: 391. (1994)

Nee.: Anales Ci. Nat. 3: 275. (1801)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus pacifica

Quercus rugosa

Quercus pacifica

Quercus rugosa