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Channel Island scrub oak, island scrub oak, Pacific oak

chêne rouge, northern red oak, red oak

Habit Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller. Trees, deciduous, to 30 m. Bark gray or dark gray, ridges wide, shiny, separated by shallow fissures, inner bark pinkish.
Bark

scaly on older branches and trunk.

Twigs

brownish or reddish, minutely puberulent, becoming glabrate and gray with age.

reddish brown, 2-3.5(-4.5) mm diam., glabrous.

Buds

light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.

Leaves

blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip;

surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.

blade ovate to elliptic or obovate, 120-200 × 60-120 mm, base broadly cuneate to almost truncate, margins with 7-11 lobes and 12-50 awns, lobes oblong, occasionally distally expanded, separated by shallow sinuses, sinuses usually extending less than 1/2 distance to midrib, apex acute;

surfaces abaxially pale green, often glaucous, glabrous except for minute axillary tufts of tomentum, adaxially dull green, glabrous, secondary veins raised on both surfaces.

Acorns

paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms;

cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed;

nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.

biennial;

cup saucer- to cup-shaped, 5-12 mm high × 18-30 mm wide, covering 1/4-1/3 nut, outer surface puberulent, inner surface light brown to red-brown, glabrous or with ring of pubescence around scar, scales less than 4 mm, often with dark margins, tips tightly appressed, obtuse;

nut ovoid to oblong, 15-30 × 10-21 mm, glabrous, scar diam. 6.5-12.5 mm.

Cotyledons

distinct.

Terminal

buds dark reddish brown, ovoid to ellipsoid, 4-7 mm, glabrous or with tuft of reddish hairs at apex.

2n

= 24.

Quercus pacifica

Quercus rubra

Phenology Flowering spring. Flowering spring.
Habitat Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands Commonly on mesic slopes and well-drained uplands, occasionally on dry slopes or poorly drained uplands
Elevation 0-300 m (0-1000 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands.

Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trees with large nuts only one-fourth covered by flat saucer-shaped cups often are treated as Quercus rubra var. rubra; those with smaller nuts one-third covered by cup- or bowl-shaped cups are treated as Q. rubra var. borealis (F. Michaux) Farwell. While E. J. Palmer (1942) suggested that these two varieties do not, K. M. McDougal and C. R. Parks (1986) found evidence of correspondence between morphologic types and flavonoid chemotypes. This is one of the most important ornamental and timber trees in the genus.

Native Americans used Quercus rubra for a number of medical purposes, including the treatment of sores, weakness, lung problems, sore throat, dysentery, indigestion, chapped skin, chills and fevers, lost voice, asthma, cough, milky urine, hear trouble, blood diseases, and Italian itch, and as an appetizer (D. E. Moerman 1986).

Quercus rubra reportedly hybridizes with Q. coccinea (= Q. ×benderi Baenitz) and Q. ellipsoidalis (P. C. Swain 1972; R. J. Jensen et al. 1993); with Q. ilicifolia (= Q. ×fernaldii Trelease), Q. imbricaria [Q. ×runcinata (A. de Candolle) Engelmann], and Q. marilandica (E. J. Palmer 1948; D. M. Hunt 1989); with Q. nigra (D. M. Hunt 1989); and with Q. palustris (= Q. ×columnaris Laughlin), Q. phellos (= Q. ×heterophylla F. Michaux), Q. shumardii (= Q. ×riparia Laughlin), and Q. velutina (= Q. ×hawkinsii Sudworth).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Lobatae
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. dumosa var. polycarpa Q. borealis, Q. maxima
Name authority Nixon & C. H. Muller: Novon 4: 391. (1994) Linnaeus: Sp. Pl. 2: 996. (1753)
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