Flora of North America species comparison

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Quercus pacifica

Quercus macrocarpa

Channel Island scrub oak, island scrub oak, Pacific oak

bur oak, burr oak, chêne à gros fruits, mossy-cup oak
Habit Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller. Trees, deciduous, to 30(-50) m. Bark dark gray, scaly or flat-ridged.
Bark

scaly on older branches and trunk.

Twigs

brownish or reddish, minutely puberulent, becoming glabrate and gray with age.

grayish or reddish, 2-4 mm diam., often forming extensive flat, radiating, corky wings, finely pubescent.
Buds

light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.

2-5(-6) mm, glabrous.
Leaves

blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip;

surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.

blade obovate to narrowly elliptic or narrowly obovate, often fiddle-shaped, (50-)70-150(-310) × (40-)50-130(-160) mm, base rounded to cuneate, margins moderately to deeply lobed, toothed, deepest sinuses near midleaf (at least in proximal 2/3), sinuses reaching nearly to midrib, longer lobes grading into shallow lobes or merely simple teeth distally, shallower, compound lobes proximally, secondary veins arched, divergent, 4-5(-10) on each side, apex broadly rounded or ovate;

surfaces abaxially light green or whitish, with minute appressed-stellate hairs forming dense, rarely sparse, tomentum, erect felty hairs absent, adaxially dark green or dull gray, sparsely puberulent to glabrate.
Acorns

paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms;

cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed;

nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.

1-3 on stout peduncle (0-)6-20(-25) mm;

cup hemispheric or turbinate, (8-)15-50 mm deep × (10-)20-60 mm wide, enclosing 1/2-7/8 nut or more, scales closely appressed, laterally connate, broadly triangular, keeled, tuberculate, finely grayish tomentose, those near margins often with soft awns to 5-10 mm or more, forming fringe around nut;

nut light brown or grayish, ovoid-ellipsoid or oblong, (15-)25-50 × (10-)20-40 mm, finely puberulent or floccose.
Cotyledons

distinct.

distinct.
2n

= 24.

Quercus pacifica

Quercus macrocarpa
Phenology Flowering spring. Flowering in spring.
Habitat Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands Bottomlands, riparian slopes, poorly drained areas, prairies, usually on limestone or calcareous clays (in nw part of range on dry slopes and ridges, prairies)
Elevation 0-300 m (0-1000 ft) 0-1000 m (0-3300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; ND; NE; NJ; NY; OH; OK; PA; SD; TN; TX; VA; VT; WI; WV; WY; MB; NB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands.

Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus macrocarpa is one of our most cold-tolerant oak species; it also endures a wide variety of other harsh conditions including poor dry soils and wet, poorly drained, and inundated locations. Putative hybrids with Q. bicolor are common in the northeastern part of its range, where the two species often occur together in wet, poorly drained habitats. The effect of this contact may be partially responsible for morphologic differences across the range of Q. macrocarpa. The large acorns are best developed in the southern part of the range, and a clinal decrease in acorn size and extent of the mossy fringe on the acorn cup seems to occur as one travels from south to north. In the northwest part of its range, Q. macrocarpa varies clinally to smaller, shrubbier forms on bluffs and hillsides, with smaller, less fringed cups, that are the basis of Q. macrocarpa var. depressa (Nuttall) Engelmann and Q. mandanensis Rydberg. These scrubby forms may merit formal recognition after more thorough study; they are treated here as clinal variants of the species. Quercus macrocarpa forms putative hybrids also with Q. alba in the savannah-type regions of the midwest. Putative hybrids with Q. gambelii occur out of the range of Q. macrocarpa.

Quercus macrocarpa is the only oak species native to Montana (in the southeast corner). Wood of Q. macrocarpa is similar to that of Q. alba and produces one of the best and most durable oak lumbers.

Native Americans used Quercus macrocarpa medicinally to treat heart troubles, cramps, diarrhea, Italian itch, and broken bones, to expel pinworms, and as an astringent (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. dumosa var. polycarpa Q. macrocarpa var. depressa, Q. mandanensis
Name authority Nixon & C. H. Muller: Novon 4: 391. (1994) Michaux: Hist. Chênes Amér., plates 2, 3. (1801)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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