FNA: Quercus pacifica vs. Quercus ajoensis

	Quercus pacifica	Quercus ajoensis
	Channel Island scrub oak, island scrub oak, Pacific oak	Ajo Mountain scrub oak
Habit	Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller.	Shrubs, rarely trees, evergreen, to 2-3 m. Bark gray, scaly or furrowed.
Bark	scaly on older branches and trunk.
Twigs	brownish or reddish, minutely puberulent, becoming glabrate and gray with age.	light brown, 1-2 mm diam., inconspicuously short stellate-pubescent or glabrate.
Buds	light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.	brown or reddish brown, ovoid or globose, 1-1.5 mm, variously short stellate-pubescent, tomentose, or glabrate.
Leaves	blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip; surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.	petiole (2-)3-4 m. Leaf blade ovate to narrowly ovate or oblong, (10-)15-35(-50) × (5-)10-20(-30) mm, rather leathery, base cordate, rarely rounded, margins crispate, sometimes flat, cartilaginous, with 4-6(-8) long-attenuate, spinose-awned teeth on each side, secondary veins 5-8 on each side, whitish, apex acute or obtuse with bristly distal teeth; surfaces abaxially blue-green, waxy-glaucous, microscopically papillose, glabrous, sometimes sparsely stellate-pubescent along midrib, adaxially blue-green, glaucous, glabrous or sparingly stellate-pubescent along midrib, secondary veins raised on both surfaces.
Acorns	paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms; cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed; nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.	solitary or paired on thin axillary peduncle (5-)30-50 mm; cup shallowly cup-shaped, thin, 3-4 mm deep × 6-8(-10) mm wide, enclosing only base of nut, scales brownish, moderately tuberculate, pubescent; nut oblong to narrowly ovoid, 12-15 × 5-8 mm.
Cotyledons	distinct.	distinct.

	Quercus pacifica	Quercus ajoensis
Phenology	Flowering spring.	Flowering in spring.
Habitat	Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands	Rare to locally abundant on igneous slopes
Elevation	0-300 m (0-1000 ft)	500-1500 m (1600-4900 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	AZ; Mexico (Baja California) [BONAP county map]
Discussion	Of conservation concern. Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands. Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Populations of Quercus ajoensis in southern New Mexico show characteristics suggesting introgression from hybridization with Quercus toumeyi, such as increased twig and leaf pubescence and sometimes the prominent golden puberulum of the abaxial leaf surfaces. Hybrids between Q. ajoensis and both Q. turbinella and Q. gambelii (Utah) are also known. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. dumosa var. polycarpa	Q. turbinella subsp. ajoensis, Q. turbinella var. ajoensis
Name authority	Nixon & C. H. Muller: Novon 4: 391. (1994)	C. H. Muller: Madroño 12: 140. (1954)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus ajoensis

Channel Island scrub oak, island scrub oak, Pacific oak

Ajo Mountain scrub oak

Habit

Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller.

Shrubs, rarely trees, evergreen, to 2-3 m. Bark gray, scaly or furrowed.

Bark

scaly on older branches and trunk.

Twigs

brownish or reddish, minutely puberulent, becoming glabrate and gray with age.

light brown, 1-2 mm diam., inconspicuously short stellate-pubescent or glabrate.

Buds

light or chestnut brown, ovate or globose, 2-3 × 1-2 mm.

brown or reddish brown, ovoid or globose, 1-1.5 mm, variously short stellate-pubescent, tomentose, or glabrate.

Leaves

blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip;

surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs.

petiole (2-)3-4 m. Leaf blade ovate to narrowly ovate or oblong, (10-)15-35(-50) × (5-)10-20(-30) mm, rather leathery, base cordate, rarely rounded, margins crispate, sometimes flat, cartilaginous, with 4-6(-8) long-attenuate, spinose-awned teeth on each side, secondary veins 5-8 on each side, whitish, apex acute or obtuse with bristly distal teeth;

surfaces abaxially blue-green, waxy-glaucous, microscopically papillose, glabrous, sometimes sparsely stellate-pubescent along midrib, adaxially blue-green, glaucous, glabrous or sparingly stellate-pubescent along midrib, secondary veins raised on both surfaces.

Acorns

paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms;

cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed;

nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate.

solitary or paired on thin axillary peduncle (5-)30-50 mm;

cup shallowly cup-shaped, thin, 3-4 mm deep × 6-8(-10) mm wide, enclosing only base of nut, scales brownish, moderately tuberculate, pubescent;

nut oblong to narrowly ovoid, 12-15 × 5-8 mm.

Cotyledons

distinct.

Quercus pacifica

Quercus ajoensis

Phenology

Flowering spring.

Flowering in spring.

Habitat

Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands

Rare to locally abundant on igneous slopes

Elevation

0-300 m (0-1000 ft)

500-1500 m (1600-4900 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

AZ; Mexico (Baja California)

[BONAP county map]

Discussion

Of conservation concern.

Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands.

Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Quercus ajoensis in southern New Mexico show characteristics suggesting introgression from hybridization with Quercus toumeyi, such as increased twig and leaf pubescence and sometimes the prominent golden puberulum of the abaxial leaf surfaces. Hybrids between Q. ajoensis and both Q. turbinella and Q. gambelii (Utah) are also known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. dumosa var. polycarpa

Q. turbinella subsp. ajoensis, Q. turbinella var. ajoensis

Name authority

Nixon & C. H. Muller: Novon 4: 391. (1994)

C. H. Muller: Madroño 12: 140. (1954)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus pacifica

Quercus ajoensis

Quercus pacifica

Quercus ajoensis