Quercus pacifica |
Quercus acerifolia |
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Channel Island scrub oak, island scrub oak, Pacific oak |
maple-leaf oak |
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Habit | Shrubs, rarely small trees, subevergreen, shrubs to 2 m, trees to 5 m or taller. | Trees or shrubs, deciduous, to 15 m. Bark dark gray to almost black, sometimes becoming rough and furrowed. |
Bark | scaly on older branches and trunk. |
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Twigs | brownish or reddish, minutely puberulent, becoming glabrate and gray with age. |
grayish brown to reddish brown, 1.5-3(-3.5) mm diam., glabrous or sparsely pubescent. |
Buds | light or chestnut brown, ovate or globose, 2-3 × 1-2 mm. |
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Leaves | blade obovate or oblong, planar to moderately convex or undulate, 15-40 × 7-20(-40) mm, base cuneate or rounded, attenuate-decurrent along petiole, margins minutely cartilaginous, entire or with 1-5 irregular teeth on each side, secondary veins obscure, 1-5 on each side, apex blunt or rounded, occasionally subacute with mucronate tip; surfaces abaxially waxy, glandular, with scattered minute, flat, appressed, ± 8-rayed stellate hairs, not obscuring surface, adaxially green, glossy, glabrate or with minute, scattered, stellate hairs. |
blade oblate to broadly elliptic, 70-140 × (60-)100-150(-180) mm, base cordate-truncate to obtuse, margins with 5-7(-9) lobes and 11-48 awns, lobes ovate-oblong or markedly distally expanded, the middle or apical pairs often overlapping, apex acute; surfaces abaxially glabrous or with prominent axillary tufts of tomentum, occasionally with scattered pubescence, adaxially glabrous, secondary veins raised on both surfaces. |
Acorns | paired or solitary in leaf axil, subsessile, rarely pedunculate in teratological forms; cup hemispheric to turbinate, to 15 mm deep × 20 mm wide, enclosing only 1/4-1/2 nut, scales moderately to heavily tuberculate, irregularly formed; nut light brown, acute-cylindric or fusiform, tapered, (15-)20-30 × (6-)9-15 m, apex acute, glabrate. |
biennial; cup saucer- to cup-shaped, 4-7 mm high × 10-20 mm wide, covering 1/4-1/3 nut, outer surface glabrous or puberulent, inner surface light brown to red-brown, glabrous or with ring of pubescence around scar, scales often with pale margins, tips tightly appressed, obtuse or acute; nut ovoid to oblong, 10.5-20 × 9-15 mm, glabrous or pubescent, scar diam. 5-9 mm. |
Cotyledons | distinct. |
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Terminal | buds gray to grayish brown, ovoid or broadly ellipsoid, 3.5-5.5 mm, glabrous. |
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Quercus pacifica |
Quercus acerifolia |
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Phenology | Flowering spring. | Flowering spring. |
Habitat | Chaparral, oak woodlands, margins of grasslands, understory in closed-cone pine stands | Dry glades, slopes, and ridge tops |
Elevation | 0-300 m (0-1000 ft) | 500-800 m (1600-2600 ft) |
Distribution |
CA
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AR |
Discussion | Of conservation concern. Quercus pacifica is endemic on three of the California Channel Islands: Santa Cruz, Santa Catalina, and Santa Rosa. It is not known from the mainland, but it bears a superficial similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations, sometimes treated as Q. ×acutidens, differ in having much greater variability in leaf shape; thicker, more leathery leaves; denser abaxial leaf vestiture; much smaller hairs, having more than 10 rays; and variable levels of connation of cotyledons (always distinct in Q. pacifica). Quercus pacifica appears to be most closely related to Q. douglasii, whether by direct descent or by introgression with another species no longer extant on the islands. Quercus ×macdonaldii Greene (as a species) [= Quercus dumosa var. macdonaldii (Greene) Jepson] is a stabilized hybrid complex between Quercus pacifica and Q. lobata Née. The plants tend to be small to moderate trees with leaves that resemble those of Q. lobata; the leaves are much more shallowly lobed and always less than two-thirds the distance from the margin to the midrib. Quercus ×macdonaldii is known from Santa Rosa, Santa Cruz, and Santa Catalina islands. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Quercus acerifolia is known only from four localities in Arkansas: Magazine Mountain, Logan County; Porter Mountain, Polk County; Pryor Mountain, Montgomery County; and Sugarloaf Mountain, Sebastian County (N. Stoynoff and W. J. Hess 1990; G. P. Johnson 1992, 1994). Some specimens suggest hybridization with Q. marilandica and/or Q. velutina, but no hybrids have been reported. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Quercus | Fagaceae > Quercus > sect. Lobatae |
Sibling taxa | ||
Synonyms | Q. dumosa var. polycarpa | Q. shumardii var. acerifolia |
Name authority | Nixon & C. H. Muller: Novon 4: 391. (1994) | (E. J. Palmer) Stoynoff & W. J. Hess ex R. J. Jensen: Fl. N. Amer. 3: 465. (1997) |
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