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chêne gris, water oak

Palmer oak, Palmer's oak

Habit Trees, deciduous or tardily deciduous, to 30 m. Bark grayish black, fissures irregular, shallow, inner bark pinkish.
Twigs

dark red-brown, 1.5-2.5 mm diam., glabrous.

Leaves

blade distinctly obtrullate, rarely elliptic or merely obovate, widest near apex, 30-120(-160) × 15-60(-70) mm, base attenuate or cuneate, rarely rounded, margins entire with 1 apical awn or with 2-3 shallow lobes and 2-5 awns (leaves on juvenile or 2d-flush growth may be deeply lobed with more awns), apex obtuse to blunt or rounded;

surfaces abaxially glabrous except for minute or conspicuous axillary tufts of tomentum, veins rarely raised, adaxially glabrous with secondary veins somewhat impressed.

blade suborbiculate, elliptic to round-ovate, 20-30(-50) × 20-40 mm, crisped, leathery and brittle, base obtuse to strongly subcordate, secondary veins 5-8(-12) pairs, each terminating in spine, basal pairs recurving, others branching at 45° angles, raised abaxially, margins spinose-dentate to occasionally entire, with highly thickened cell walls, spines cartilaginous, (1-)1.5-2 mm, apex broadly rounded or subacute, spinose;

surfaces abaxially glaucous with waxy layer, often obscured by golden brown glandular hairs, adaxially grayish dark green, scurfy with fasciculate erect and twisting hairs.

Acorns

biennial;

cup saucer-shaped, 2.5-5.5 mm high × 10-18 mm wide, covering 1/4 nut or less, outer surface puberulent, inner surface sparsely to uniformly pubescent, scale tips tightly appressed, acute;

nut broadly ovoid, 9.5-14 × 9.5-14.5 mm, often faintly striate, glabrate, scar diam. 6-11.5 mm.

solitary or rarely paired;

cup turbinate to saucer-shaped, margins involute, often irregular, 7-10 mm deep × 10-25(-35) mm wide, scales appressed, embedded, often appearing laterally connate into concentric rings with only tip of scale visible, tuberculate, densely golden-tomentose throughout;

nut oblong to fusiform, 20-30 × 10-15 mm, apex acute.

Terminal

buds reddish brown, ovoid, 3-6.5 mm, puberulent throughout, occasionally densely pubescent on apical 2/3.

buds ovoid, 1-1.5 mm, apex rounded, glabrous.

Small

trees and shrubs, to 2-3 m. Twigs rigid, divaricately branched at 65-90° angles, reddish brown, 1.5-3 mm diam., pubescent, sparsely so in 2d year.

Quercus nigra

Quercus palmeri

Phenology Flowering spring. Flowering in spring.
Habitat Mesic alluvial and lowland sites, also barrens, dunes, hammocks, and low ridges to steep slopes Disjunct in canyons, mountain washes, dry thickets, and margins of chapparal communities
Elevation 0-450 m (0-1500 ft) 700-1800 m (2300-5900 ft)
Distribution
from FNA
AL; AR; DE; FL; GA; KY; LA; MD; MO; MS; NC; NJ; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; Mexico (n Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Typically on mesic alluvial and lowland sites, Quercus nigra also occurs on a wide variety of soil types and in a diversity of habitats.

Trees with 3-lobed leaves with attenuate bases have been recognized as Quercus nigra var. tridentifera Sargent.

Quercus nigra reportedly hybridizes with Q. falcata (= Q. ×garlandensis E. J. Palmer), Q. incana, Q. laevis (= Q. ×walteriana Ashe), Q. marilandica (= Q. ×sterilis Trelease), Q. phellos (= Q. ×capesii W. Wolf), Q. shumardii (= Q. ×neopalmeri Sudworth), and Q. velutina (Q. ×demarei Ashe). In addition, D. M. Hunt (1989) cited evidence of hybridization also with Q. arkansana, Q. georgiana, Q. hemisphaerica, Q. laurifolia, Q. myrtifolia, Q. palustris, Q. rubra, and Q. texana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Quercus palmeri are often small and may exist as single clones. The disjunct populations of California and Baja California are consistent morphologically. In Arizona populations, individuals tend to have flatter leaves bearing fewer teeth; this distinction is not constant, however. Morphologically aberrant populations identified as Q. palmeri in eastern Arizona (Chiracahua, Huachuca, and Santa Catalina mountains) and southwestern New Mexico are most likely the result of introgression from Q. palmeri to Q. chrysolepis (J. M. Tucker and H. S. Haskell 1960). Those populations tend to be intermediate in overall morphology, but all lack the diagnostic trichomes and biochemical markers of Q. palmeri; they are best classified as Q. chrysolepis affinity Q. palmeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Lobatae Fagaceae > Quercus > sect. Protobalanus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. nana, Q. nigra var. tridentifera, Q. uliginosa Q. dunnii
Name authority Linnaeus: Sp. Pl. 2: 995. (1753) Engelmann: in S. Watson, Bot. California 2: 97. (1880)
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