Quercus nigra |
Quercus havardii |
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chêne gris, water oak |
Havard oak, Havard's oak, Havard's shinnery oak, sand shinnery oak, sand shinoak, shinnery oak |
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Habit | Trees, deciduous or tardily deciduous, to 30 m. Bark grayish black, fissures irregular, shallow, inner bark pinkish. | Shrubs, deciduous, low, forming clones 0.3-1.5 × 10 m, rhizomatous. |
Bark | light gray, scaly-papery. |
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Twigs | dark red-brown, 1.5-2.5 mm diam., glabrous. |
brown or grayish, 1-2.5 mm diam., glabrous or densely short grayish or yellowish tomentulose, glabrate in age. |
Buds | dark red-brown, subglobose, ca. 2 mm, sparsely pubescent. |
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Leaves | blade distinctly obtrullate, rarely elliptic or merely obovate, widest near apex, 30-120(-160) × 15-60(-70) mm, base attenuate or cuneate, rarely rounded, margins entire with 1 apical awn or with 2-3 shallow lobes and 2-5 awns (leaves on juvenile or 2d-flush growth may be deeply lobed with more awns), apex obtuse to blunt or rounded; surfaces abaxially glabrous except for minute or conspicuous axillary tufts of tomentum, veins rarely raised, adaxially glabrous with secondary veins somewhat impressed. |
blade green, often turning brownish with age, polymorphic, oblong or elliptic or sometimes lanceolate to oblanceolate or ovate to obovate, (30-)50-100 × (10-)20-50 mm, rather thick and hard, base rounded to cuneate, margins flat to revolute, at least some undulate, 2-3 rounded teeth on each side, secondary veins 5-8 on each side, much branched, apex broadly rounded, rarely acute; surfaces abaxially densely grayish or yellowish tomentulose or stellate-pubescent, sometimes only sparsely pubescent, secondary veins quite prominent, adaxially lustrous, very sparsely stellate-pubescent or glabrate, secondary veins very slightly if at all raised. |
Acorns | biennial; cup saucer-shaped, 2.5-5.5 mm high × 10-18 mm wide, covering 1/4 nut or less, outer surface puberulent, inner surface sparsely to uniformly pubescent, scale tips tightly appressed, acute; nut broadly ovoid, 9.5-14 × 9.5-14.5 mm, often faintly striate, glabrate, scar diam. 6-11.5 mm. |
solitary or paired, subsessile or on peduncle to 10(-18) mm; cup from deeply cup-shaped to goblet-shaped, 10-12 mm deep × 15-25 mm wide, enclosing 1/3-1/2 nut, base rounded or slightly constricted, margin very thin and smooth, scales reddish brown, triangular-ovate to long-acute, proximally moderately to markedly tuberculate, pubescent, often canescent, tips loosely appressed; nut brown, ovoid, 12-25 × 14-18 mm. |
Cotyledons | distinct. |
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Terminal | buds reddish brown, ovoid, 3-6.5 mm, puberulent throughout, occasionally densely pubescent on apical 2/3. |
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Quercus nigra |
Quercus havardii |
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Phenology | Flowering spring. | Flowering spring. |
Habitat | Mesic alluvial and lowland sites, also barrens, dunes, hammocks, and low ridges to steep slopes | Deep, shifting or stabilized sand dunes, off deep sands in putative hybrid populations |
Elevation | 0-450 m (0-1500 ft) | 500-1500 m (1600-4900 ft) |
Distribution |
AL; AR; DE; FL; GA; KY; LA; MD; MO; MS; NC; NJ; OK; SC; TN; TX; VA
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NM; OK; TX
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Discussion | Typically on mesic alluvial and lowland sites, Quercus nigra also occurs on a wide variety of soil types and in a diversity of habitats. Trees with 3-lobed leaves with attenuate bases have been recognized as Quercus nigra var. tridentifera Sargent. Quercus nigra reportedly hybridizes with Q. falcata (= Q. ×garlandensis E. J. Palmer), Q. incana, Q. laevis (= Q. ×walteriana Ashe), Q. marilandica (= Q. ×sterilis Trelease), Q. phellos (= Q. ×capesii W. Wolf), Q. shumardii (= Q. ×neopalmeri Sudworth), and Q. velutina (Q. ×demarei Ashe). In addition, D. M. Hunt (1989) cited evidence of hybridization also with Q. arkansana, Q. georgiana, Q. hemisphaerica, Q. laurifolia, Q. myrtifolia, Q. palustris, Q. rubra, and Q. texana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Individual clones emerging to heights of 2-3 m from thickets occur sporadically across the Texas range of Quercus havardii and express some characteristics of Q. stellata, such as more deeply lobed leaves and smaller acorns. Such putative hybrids increase in frequency in the eastern part of the range of the species. Material of Quercus havardii from the Navajo Basin of Utah and adjacent Arizona has been treated as Q. havardii var. tuckeri Welsh. Welsh followed J. M. Tucker (1970) and interpreted these intermediate populations as putative hybrids between Q. havardii and both Q. turbinella and Q. gambelii. Giving varietal rank, instead of nothospecies status, to such populations seems arbitrary, and it certainly is inconsistent with their putative hybrid origins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Lobatae | Fagaceae > Quercus > sect. Quercus |
Sibling taxa | ||
Synonyms | Q. nana, Q. nigra var. tridentifera, Q. uliginosa | |
Name authority | Linnaeus: Sp. Pl. 2: 995. (1753) | Rydberg: Bull. New York Bot. Gard. 2: 213. (1901) |
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