FNA: Quercus john-tuckeri vs. Quercus pungens

	Quercus john-tuckeri	Quercus pungens
	desert scrub oak, Tucker oak, Tucker's oak	pungent oak, sandpaper oak
Habit	Shrubs, subevergreen or evergreen, 1-3(-5) m. Bark light gray or brown, scaly.	Shrubs or moderate-sized trees, evergreen or subevergreen.
Bark		light brown, papery.
Twigs	yellowish or dingy gray, 1-1.5(-2) mm diam., densely tomentulose.	gray, 1-2 mm diam., short velvety-tomentose, glabrate with age.
Buds	brown, ovoid or globose, 1.5-2(-3) mm, glabrous except for ciliate margins of scales; proximal scales often yellowish puberulent.	dark red-brown, ca. 2 mm, sparsely pubescent.
Leaves	blade unicolored, elliptic or obovate, (10-)15-30(-40) × (8-)10-15(-20) mm, thick and leathery, often brittle, base truncate or rounded-attenuate, rarely subcordate, margins irregularly spinose-toothed, occasionally shallowly lobate, secondary veins (3-)4-7, often some veins branching near margin and passing into more than 1 tooth, apex acute or rounded; surfaces abaxially waxy grayish, light green, or yellowish, sparse to moderately dense (8-)10-12-rayed, (loosely) appressed-stellate hairs, often 0.2-0.5 mm diam., and sparse to crowded, yellowish, glandular hairs, adaxially dull grayish, with stellate hairs, similar to abaxial surface.	blade elliptic to oblong, 10-40(-90) × 10-20(-40) mm, rather thick, leathery, stiff, base rounded or minutely cordate, very rarely cuneate, margins regularly undulate-crisped, not revolute, coarsely toothed or incised with acute teeth or spinose lobes, secondary veins 5-8(-14) on each side, usually branched before passing into teeth, apex acute or obtuse, rarely rounded, spine-tipped; surfaces abaxially canescent, usually densely stellate-pubescent, and mixed with stiff, harsh, stellate hairs, often sandpapery to touch, rarely glabrate, adaxially yellowish green, glossy, usually rough and sandpapery because of minute, persistent hair bases, rarely glabrate.
Acorns	solitary or paired, subsessile; cup cup-shaped or obconic to hemispheric, 5-7 mm deep × 10-15 mm wide, thin, scales whitish or yellowish, moderately or scarcely tuberculate, puberulent; nut fusiform, ovoid, or conic, 20-30 mm, apex acute.	subsessile or on peduncle to 3 mm; cup shallowly to deeply cup-shaped or turbinate, to 8 mm deep × 13 mm wide, covering ca. 1/4 nut, margin thin, scales reddish brown, moderately tuberculate or keeled, densely gray-tomentose; nut light brown, broadly ovoid to subcylindric, to 10 × 10 mm, apex rounded to subacute, glabrous.
Cotyledons	distinct.	distinct.

	Quercus john-tuckeri	Quercus pungens
Phenology		Flowering spring.
Habitat	Dry slopes, chaparral, pinyon and juniper woodlands, margins of oak woodlands and sagebrush	On dry limestone or igneous slopes, usually in oak, pinyon, and juniper woodlands, chaparral, and sometimes descending into desert vegetation
Elevation	900-2000 m (3000-6600 ft)	800-2000 m (2600-6600 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	AZ; NM; TX; Mexico (Chihuahua and Coahuila) [WildflowerSearch map] [BONAP county map]
Discussion	Dry slopes, chaparral, pinyon and juniper woodlands, margins of oak woodlands and sagebrush; 900-2000 m; Calif. Endemic to California, Quercus john-tuckeri occurs from Los Angeles County northward in the interior Coast Ranges and Sierra Foothills to the northern edge of Sacramento Valley. Quercus john-tuckeri bears some resemblance to both Q. turbinella and Q. berberidifolia. Quercus turbinella has pedunculate fruit and cordate leaf bases, however, and Q. berberidifolia has a glabrate adaxial leaf surface, substantially smaller stellate trichomes with fewer rays on the abaxial leaf surface, heavier tuberculate acorn cups, and blunt or rounded (instead of acute) acorns. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Numerous populations appear to be hybrid swarms between Quercus pungens and Q. vaseyana, which is sometimes treated as a variety of Q. pungens. No other evidence for a close relationship exists for these two species, and such a treatment risks erecting a polyphyletic assemblage. To the west and south within the range of Q. pungens no indication of introgression exists, and the two species are strikingly different and easily separable. I interpret the contact as secondary. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. turbinella subsp. californica	Q. undulata var. pungens
Name authority	Nixon & C. H. Muller: Novon 4: 391. (1994)	Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171. 185: not Q. pungens Gandoger 1890
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus pungens

desert scrub oak, Tucker oak, Tucker's oak

pungent oak, sandpaper oak

Habit

Shrubs, subevergreen or evergreen, 1-3(-5) m. Bark light gray or brown, scaly.

Shrubs or moderate-sized trees, evergreen or subevergreen.

Bark

light brown, papery.

Twigs

yellowish or dingy gray, 1-1.5(-2) mm diam., densely tomentulose.

gray, 1-2 mm diam., short velvety-tomentose, glabrate with age.

Buds

brown, ovoid or globose, 1.5-2(-3) mm, glabrous except for ciliate margins of scales;

proximal scales often yellowish puberulent.

dark red-brown, ca. 2 mm, sparsely pubescent.

Leaves

blade unicolored, elliptic or obovate, (10-)15-30(-40) × (8-)10-15(-20) mm, thick and leathery, often brittle, base truncate or rounded-attenuate, rarely subcordate, margins irregularly spinose-toothed, occasionally shallowly lobate, secondary veins (3-)4-7, often some veins branching near margin and passing into more than 1 tooth, apex acute or rounded;

surfaces abaxially waxy grayish, light green, or yellowish, sparse to moderately dense (8-)10-12-rayed, (loosely) appressed-stellate hairs, often 0.2-0.5 mm diam., and sparse to crowded, yellowish, glandular hairs, adaxially dull grayish, with stellate hairs, similar to abaxial surface.

blade elliptic to oblong, 10-40(-90) × 10-20(-40) mm, rather thick, leathery, stiff, base rounded or minutely cordate, very rarely cuneate, margins regularly undulate-crisped, not revolute, coarsely toothed or incised with acute teeth or spinose lobes, secondary veins 5-8(-14) on each side, usually branched before passing into teeth, apex acute or obtuse, rarely rounded, spine-tipped;

surfaces abaxially canescent, usually densely stellate-pubescent, and mixed with stiff, harsh, stellate hairs, often sandpapery to touch, rarely glabrate, adaxially yellowish green, glossy, usually rough and sandpapery because of minute, persistent hair bases, rarely glabrate.

Acorns

solitary or paired, subsessile;

cup cup-shaped or obconic to hemispheric, 5-7 mm deep × 10-15 mm wide, thin, scales whitish or yellowish, moderately or scarcely tuberculate, puberulent;

nut fusiform, ovoid, or conic, 20-30 mm, apex acute.

subsessile or on peduncle to 3 mm;

cup shallowly to deeply cup-shaped or turbinate, to 8 mm deep × 13 mm wide, covering ca. 1/4 nut, margin thin, scales reddish brown, moderately tuberculate or keeled, densely gray-tomentose;

nut light brown, broadly ovoid to subcylindric, to 10 × 10 mm, apex rounded to subacute, glabrous.

Cotyledons

distinct.

Quercus john-tuckeri

Quercus pungens

Phenology

Flowering spring.

Habitat

Dry slopes, chaparral, pinyon and juniper woodlands, margins of oak woodlands and sagebrush

On dry limestone or igneous slopes, usually in oak, pinyon, and juniper woodlands, chaparral, and sometimes descending into desert vegetation

Elevation

900-2000 m (3000-6600 ft)

800-2000 m (2600-6600 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

AZ; NM; TX; Mexico (Chihuahua and Coahuila)

[WildflowerSearch map]

[BONAP county map]

Discussion

Dry slopes, chaparral, pinyon and juniper woodlands, margins of oak woodlands and sagebrush; 900-2000 m; Calif.

Endemic to California, Quercus john-tuckeri occurs from Los Angeles County northward in the interior Coast Ranges and Sierra Foothills to the northern edge of Sacramento Valley.

Quercus john-tuckeri bears some resemblance to both Q. turbinella and Q. berberidifolia. Quercus turbinella has pedunculate fruit and cordate leaf bases, however, and Q. berberidifolia has a glabrate adaxial leaf surface, substantially smaller stellate trichomes with fewer rays on the abaxial leaf surface, heavier tuberculate acorn cups, and blunt or rounded (instead of acute) acorns.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Numerous populations appear to be hybrid swarms between Quercus pungens and Q. vaseyana, which is sometimes treated as a variety of Q. pungens. No other evidence for a close relationship exists for these two species, and such a treatment risks erecting a polyphyletic assemblage. To the west and south within the range of Q. pungens no indication of introgression exists, and the two species are strikingly different and easily separable. I interpret the contact as secondary.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. turbinella subsp. californica

Q. undulata var. pungens

Name authority

Nixon & C. H. Muller: Novon 4: 391. (1994)

Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171. 185: not Q. pungens Gandoger 1890

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus john-tuckeri

Quercus pungens

Quercus john-tuckeri

Quercus pungens