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escarpment live oak, live oak, plateau oak, Texas live oak

bastard oak, bastard white oak, Durand oak, Durand white oak

Habit Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands. Trees or shrubs, deciduous, to 15(-20) m, with solitary or multiple trunks.
Bark

dark brown or black, scaly.

gray to light brown, flaky to papery and exfoliating.

Twigs

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

light gray or gray, 1-2(-3) mm diam., glabrous, rarely minutely puberulent.

Buds

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

brown or reddish brown, broadly ovoid, 2-3 mm, essentially glabrous.

Leaves

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

blade oblong to oblanceolate, or narrowly rhomboid, or cuneiform, or rounded-3-dentate, (25-)30-120(-140) × (15-)25-60 mm, base acute, cuneate, attenuate-rounded, or obtuse, margins entire to irregularly toothed or moderately, sinuately lobed, flat, secondary veins ca. 7-11 on each side, apex broadly rounded, rarely attenuately narrowed or obscurely 3-lobed;

surfaces abaxially silvery or dull green, with scattered to crowded, minute, appressed-stellate, 8-10-rayed hairs, or glabrate or glabrous, especially in shade forms, adaxially green or dull green, glabrous.

Acorns

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

solitary or paired, subsessile or on axillary peduncle to 1-7 mm;

cup saucer-shaped to shallowly cup-shaped, rarely deeper, 2-8 mm deep × 8-15(-20) mm wide, enclosing 1/8-1/4 nut, rarely more, base flat, rounded, or constricted, margin thin, scales closely appressed, grayish with reddish margins, ovate, flat, obtuse, not tuberculate;

nut light brown, depressed-ovoid to oblong, 7-15 × 7-12(-17) mm, glabrous.

Cotyledons

connate.

distinct.

Quercus fusiformis

Quercus sinuata

Phenology Flowering spring.
Habitat Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel
Elevation 0-1200 m (0-3900 ft)
Distribution
from FNA
OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The question of the correct name for this species has persisted, with some authors rejecting the usage here in favor of Quercus durandii. Although no type material is extant, the original description of Q. sinuata is consistent with the concept presented here, as by W. W. Ashe (1916) and W. Trelease (1924), and inconsistent with any other oak from the broad area covered by Thomas Walter's Flora Caroliniana (1788).

The two varieties differ in habit, habitat, leaf size and lobing, and geographic range, and considerable variability exists within both varities as to the degree and density of silvery stellate-pubescence on the abaxial surface of the leaf. Sun leaves of both tend to have a higher proportion of silvery pubescence, and shade leaves and some individual trees tend to have more glabrate leaves, although evidence of flat-stellate trichomes is usually apparent. Plants with young, expanding leaves sometimes are mistaken for Quercus nigra, a member of the red oak group.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trees to 20 m, trunk usually solitary; moist bottomlands and riparian habitats.
var. sinuata
1. Shrubs or small trees to 3(–5) m, often forming thickets, trunks multiple from near ground; dry limestone hills.
var. breviloba
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Subordinate taxa
Q. sinuata var. breviloba, Q. sinuata var. sinuata
Synonyms Q. virginiana var. fusiformis Q. durandii
Name authority Small: Bull. Torrey Bot. Club 28: 357. (1901) Walter: Fl. Carol., 235. (1788)
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