Flora of North America species comparison

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Quercus fusiformis

Quercus shumardii

escarpment live oak, live oak, plateau oak, Texas live oak

schneck oak, Shumard oak, Shumard red oak, Shumard's oak, swamp red oak
Habit Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands. Trees, deciduous, to 35 m. Bark gray-brown to dark brown, shallowly fissured with scaly or light-colored flat ridges, inner bark pinkish.
Bark

dark brown or black, scaly.

Twigs

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

gray to light brown, (1.5-)2-3.5(-4.5) mm diam., glabrous.
Buds

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

Leaves

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

blade broadly elliptic to obovate, 100-200 × 60-150 mm, base obtuse to truncate, occasionally acute, margins with 5-9 lobes and 15-50 awns, lobes oblong or distally expanded, apex acute;

surfaces abaxially glabrous except for prominent axillary tufts of tomentum, adaxially glossy, glabrous, secondary veins raised on both surfaces.
Acorns

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

biennial;

cup saucer-shaped to cup-shaped, 7-12 mm high × 15-30 mm wide, covering 1/4-1/3 nut, outer surface glabrous or puberulent, inner surface light-brown to red-brown, glabrous or with ring of pubescence around scar, scales often with pale margins, tips tightly appressed, obtuse or acute;

nut ovoid to oblong, occasionally subglobose, 14-30 × 10-20 mm, glabrous, scar diam. 6.5-12 mm.
Cotyledons

connate.

Terminal

buds gray to grayish brown, ovoid or broadly ellipsoid, 4-8 mm, often noticeably 5-angled in cross section, glabrous.

Quercus fusiformis

Quercus shumardii
Phenology Flowering spring. Flowering spring.
Habitat Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel Mesic slopes and bottoms, stream banks and poorly drained uplands
Elevation 0-1200 m (0-3900 ft) 0-500 m (0-1600 ft)
Distribution
from FNA
OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NE; OH; OK; PA; SC; TN; TX; VA; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trees with shallow cups covering ca. one-fourth of the nut are treated as Quercus shumardii var. shumardii; those with more deeply rounded cups covering ca. one-third of the nut are treated as Q. shumardii var. schneckii (Britton) Sargent. Quercus shumardii var. stenocarpa Laughlin was described from several trees in Missouri and Illinois having ellipsoid acorns that were covered less than one-third their length by very small (5.5-7 mm high × 12.5-18 mm wide), shallow cups (K. Laughlin 1969).

Quercus shumardii reportedly hybridizes with Q. buckleyi, Q. falcata (= Q. ×joori Trelease), Q. hemisphaerica, Q. imbricaria (= Q. ×egglestoni Trelease), Q. laevis, Q. laurifolia, Q. marilandica, Q. nigra, Q. palustris (= Q. ×mutabilis E. J. Palmer & Steyermark), Q. phellos (= Q. ×moultonensis Ashe), Q. rubra, and Q. velutina (= Q. × discreta Laughlin).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Lobatae
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. virginiana var. fusiformis Q. schneckii
Name authority Small: Bull. Torrey Bot. Club 28: 357. (1901) Buckley: Proc. Acad. Nat. Sci. Philadelphia 12: 444. (1860)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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