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escarpment live oak, live oak, plateau oak, Texas live oak

chêne rouge, northern red oak, red oak

Habit Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands. Trees, deciduous, to 30 m. Bark gray or dark gray, ridges wide, shiny, separated by shallow fissures, inner bark pinkish.
Bark

dark brown or black, scaly.

Twigs

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

reddish brown, 2-3.5(-4.5) mm diam., glabrous.

Buds

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

Leaves

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

blade ovate to elliptic or obovate, 120-200 × 60-120 mm, base broadly cuneate to almost truncate, margins with 7-11 lobes and 12-50 awns, lobes oblong, occasionally distally expanded, separated by shallow sinuses, sinuses usually extending less than 1/2 distance to midrib, apex acute;

surfaces abaxially pale green, often glaucous, glabrous except for minute axillary tufts of tomentum, adaxially dull green, glabrous, secondary veins raised on both surfaces.

Acorns

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

biennial;

cup saucer- to cup-shaped, 5-12 mm high × 18-30 mm wide, covering 1/4-1/3 nut, outer surface puberulent, inner surface light brown to red-brown, glabrous or with ring of pubescence around scar, scales less than 4 mm, often with dark margins, tips tightly appressed, obtuse;

nut ovoid to oblong, 15-30 × 10-21 mm, glabrous, scar diam. 6.5-12.5 mm.

Cotyledons

connate.

Terminal

buds dark reddish brown, ovoid to ellipsoid, 4-7 mm, glabrous or with tuft of reddish hairs at apex.

2n

= 24.

Quercus fusiformis

Quercus rubra

Phenology Flowering spring. Flowering spring.
Habitat Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel Commonly on mesic slopes and well-drained uplands, occasionally on dry slopes or poorly drained uplands
Elevation 0-1200 m (0-3900 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trees with large nuts only one-fourth covered by flat saucer-shaped cups often are treated as Quercus rubra var. rubra; those with smaller nuts one-third covered by cup- or bowl-shaped cups are treated as Q. rubra var. borealis (F. Michaux) Farwell. While E. J. Palmer (1942) suggested that these two varieties do not, K. M. McDougal and C. R. Parks (1986) found evidence of correspondence between morphologic types and flavonoid chemotypes. This is one of the most important ornamental and timber trees in the genus.

Native Americans used Quercus rubra for a number of medical purposes, including the treatment of sores, weakness, lung problems, sore throat, dysentery, indigestion, chapped skin, chills and fevers, lost voice, asthma, cough, milky urine, hear trouble, blood diseases, and Italian itch, and as an appetizer (D. E. Moerman 1986).

Quercus rubra reportedly hybridizes with Q. coccinea (= Q. ×benderi Baenitz) and Q. ellipsoidalis (P. C. Swain 1972; R. J. Jensen et al. 1993); with Q. ilicifolia (= Q. ×fernaldii Trelease), Q. imbricaria [Q. ×runcinata (A. de Candolle) Engelmann], and Q. marilandica (E. J. Palmer 1948; D. M. Hunt 1989); with Q. nigra (D. M. Hunt 1989); and with Q. palustris (= Q. ×columnaris Laughlin), Q. phellos (= Q. ×heterophylla F. Michaux), Q. shumardii (= Q. ×riparia Laughlin), and Q. velutina (= Q. ×hawkinsii Sudworth).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Lobatae
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. virginiana var. fusiformis Q. borealis, Q. maxima
Name authority Small: Bull. Torrey Bot. Club 28: 357. (1901) Linnaeus: Sp. Pl. 2: 996. (1753)
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