FNA: Quercus fusiformis vs. Quercus hemisphaerica

	Quercus fusiformis	Quercus hemisphaerica
	escarpment live oak, live oak, plateau oak, Texas live oak	Darlington oak, Darlington's oak, laurel oak, sand laurel oak
Habit	Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands.	Trees, evergreen or tardily deciduous, to 35 m. Bark much like that of Q. laurifolia.
Bark	dark brown or black, scaly.
Twigs	light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.	light brown to dark red-brown, 1-2.5 mm diam., glabrous.
Buds	reddish or dark brown, subglobose or ovate, 1.5-3 mm; scale margins glabrous or puberulent.
Leaves	blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute; surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.	blade narrowly ovate or elliptic to oblanceolate, 30-120 × 10-40 mm, leathery, base obtuse to rounded, rarely attenuate, margins entire or with a few shallow lobes or teeth near apex, awns 1-4 (rarely as many as 8-10 on 2d-flush growth), apex acute or acuminate, occasionally obtuse; surfaces abaxially glabrous, rarely with minute axillary tufts of tomentum, adaxially glabrous.
Acorns	1-3, on peduncle 3-30 mm; cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent; nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.	biennial; cup saucer-shaped to bowl-shaped, rarely turbinate, 3-10 mm high × 11-18 mm wide, covering 1/4-1/3 nut, outer surface puberulent, inner surface pubescent at least 1/2 distance to rim, scales occasionally distinctly tuberculate, tips appressed, acute to obtuse; nut broadly ovoid to hemispheric, 9-16 × 9-16 mm, glabrate, scar diam. 6-9.5 mm.
Cotyledons	connate.
Terminal		buds reddish to purplish brown, ovoid, 2.5-5 mm, glabrous or with ciliate scale margins.

	Quercus fusiformis	Quercus hemisphaerica
Phenology	Flowering spring.	Flowering spring.
Habitat	Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel	Moderately dry sandy soils, scrub sandhills, stream banks, occasionally on hillsides and ravines
Elevation	0-1200 m (0-3900 ft)	0-150 m (0-500 ft)
Distribution	OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas) [WildflowerSearch map] [BONAP county map]	AL; AR; FL; GA; LA; MS; NC; SC; TX; VA [WildflowerSearch map] [BONAP county map]
Discussion	The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns. Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Quercus hemisphaerica flowers about two weeks later than sympatric Q. laurifolia (W. H. Duncan and M. B. Duncan 1988). Most authors have treated Quercus hemisphaerica as synonymous with Q. laurifolia. M. L. Fernald (1946) carefully examined the situation and concluded that Q. hemisphaerica is a distinct entity, but C. H. Muller (1951) argued that these two taxa "...are now certainly not separable even as varieties of the same species." Later (1970), Muller recanted by recognizing Q. hemisphaerica as a common component of stream terraces along the Gulf Coast. Quercus hemisphaerica reportedly hybridizes with Q. falcata (C. H. Muller 1970); with Q. arkansana, Q. inopina, Q. marilandica, Q. myrtifolia, Q. nigra, Q. phellos, Q. pumila, and Q. shumardii (D. M. Hunt 1989); and with Q. incana (producing Q. ×sublaurifolia Trelease), and Q. laevis (producing Q. ×mellichampi Trelease). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Lobatae
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. virginiana var. fusiformis
Name authority	Small: Bull. Torrey Bot. Club 28: 357. (1901)	W. Bartram ex Willdenow: Sp. Pl. 4(1): 443. (1805)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus hemisphaerica

escarpment live oak, live oak, plateau oak, Texas live oak

Darlington oak, Darlington's oak, laurel oak, sand laurel oak

Habit

Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands.

Trees, evergreen or tardily deciduous, to 35 m. Bark much like that of Q. laurifolia.

Bark

dark brown or black, scaly.

Twigs

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

light brown to dark red-brown, 1-2.5 mm diam., glabrous.

Buds

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

Leaves

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

blade narrowly ovate or elliptic to oblanceolate, 30-120 × 10-40 mm, leathery, base obtuse to rounded, rarely attenuate, margins entire or with a few shallow lobes or teeth near apex, awns 1-4 (rarely as many as 8-10 on 2d-flush growth), apex acute or acuminate, occasionally obtuse;

surfaces abaxially glabrous, rarely with minute axillary tufts of tomentum, adaxially glabrous.

Acorns

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

biennial;

cup saucer-shaped to bowl-shaped, rarely turbinate, 3-10 mm high × 11-18 mm wide, covering 1/4-1/3 nut, outer surface puberulent, inner surface pubescent at least 1/2 distance to rim, scales occasionally distinctly tuberculate, tips appressed, acute to obtuse;

nut broadly ovoid to hemispheric, 9-16 × 9-16 mm, glabrate, scar diam. 6-9.5 mm.

Cotyledons

connate.

Terminal

buds reddish to purplish brown, ovoid, 2.5-5 mm, glabrous or with ciliate scale margins.

Quercus fusiformis

Quercus hemisphaerica

Phenology

Flowering spring.

Habitat

Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel

Moderately dry sandy soils, scrub sandhills, stream banks, occasionally on hillsides and ravines

Elevation

0-1200 m (0-3900 ft)

0-150 m (0-500 ft)

Distribution

OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)

[WildflowerSearch map]

[BONAP county map]

AL; AR; FL; GA; LA; MS; NC; SC; TX; VA

[WildflowerSearch map]

[BONAP county map]

Discussion

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus hemisphaerica flowers about two weeks later than sympatric Q. laurifolia (W. H. Duncan and M. B. Duncan 1988).

Most authors have treated Quercus hemisphaerica as synonymous with Q. laurifolia. M. L. Fernald (1946) carefully examined the situation and concluded that Q. hemisphaerica is a distinct entity, but C. H. Muller (1951) argued that these two taxa "...are now certainly not separable even as varieties of the same species." Later (1970), Muller recanted by recognizing Q. hemisphaerica as a common component of stream terraces along the Gulf Coast.

Quercus hemisphaerica reportedly hybridizes with Q. falcata (C. H. Muller 1970); with Q. arkansana, Q. inopina, Q. marilandica, Q. myrtifolia, Q. nigra, Q. phellos, Q. pumila, and Q. shumardii (D. M. Hunt 1989); and with Q. incana (producing Q. ×sublaurifolia Trelease), and Q. laevis (producing Q. ×mellichampi Trelease).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Fagaceae > Quercus > sect. Lobatae

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. virginiana var. fusiformis

Name authority

Small: Bull. Torrey Bot. Club 28: 357. (1901)

W. Bartram ex Willdenow: Sp. Pl. 4(1): 443. (1805)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Quercus fusiformis

Quercus hemisphaerica

Quercus fusiformis

Quercus hemisphaerica