Flora of North America species comparison

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Quercus fusiformis

Quercus chrysolepis

escarpment live oak, live oak, plateau oak, Texas live oak

canyon live oak, gold cup live oak, maul oak
Habit Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands. Trees or shrubs, trees small to medium-sized, to 25 m, shrubs of variable size.
Bark

dark brown or black, scaly.

Twigs

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

branching at 60° angles or less, golden brown, 1-2 mm diam., flexible, densely pubescent 1st year, moderately so 2d year.
Buds

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

Leaves

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

blade oblong, acuminate, usually flat to slightly concave, 20-70 × 10-35 mm, thick, leathery, base obtuse to rounded, secondary veins 12 or more pairs, branching at ca. 50° angles, slightly raised abaxially, margins often slightly revolute with moderately thickened cell walls, entire to spinulose-dentate (especially on juvenile growth), regularly toothed, teeth terminating with mucronate to spinescent tip, apex acute or obtuse, mucronate to spinescent;

surfaces abaxially glabrate to pubescent with bluish white wax layer, often obscured by golden glandular and multiradiate hairs, adaxially yellowish green, scurfy with multiradiate hairs, later in season slightly pubescent.
Acorns

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

solitary or paired, rarely in 3s or 4s;

cup saucer-shaped, 4-10 mm deep × 15-40 mm wide, rims often corky and thickened, scales appressed, deeply embedded in tomentum, often appearing swollen and keeled, tuberculate;

nut ovoid, 15-30 × 10-20 mm, apex blunt, glabrous;

nut scar 4-10 mm diam.
Cotyledons

connate.

Terminal

buds conic, 2-8 mm, scales brown with ciliate margins.

Quercus fusiformis

Quercus chrysolepis
Phenology Flowering spring. Flowering usually in spring, occasionally in fall.
Habitat Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel Mountain ridges, canyons, and moist slopes
Elevation 0-1200 m (0-3900 ft) 200-2600 m (700-8500 ft)
Distribution
from FNA
OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; OR; Mexico (Baja California and Chihuahua)
[WildflowerSearch map]
[BONAP county map]
Discussion

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus chrysolepis is one of the most variable North American oaks. Historically, individuals with extreme variation in fruit and leaf characteristics led to the recognition of several varieties and forms; for example, shrubs with small leaves have been called Q. chrysolepis var. nana (Jepson) Jepson. Studies of quantitative and qualitative variation in these characteristics do not support the recognition of infraspecific taxa; geographic variation is apparent, however, based on populations with nearly stabilized character combinations that loosely define widespread variants. This oak is distinguished consistently from other species of the complex by the presence of multiradiate trichomes on both leaf surfaces (P. S. Manos 1993). The leaf morphology and branching habit present on juvenile growth, suckers, and shade forms may approach typical Q. palmeri. Similarly, various extreme forms often resemble other species of the complex.

Putative hybrids have been reported from narrow zones of range overlap with the three other North American species of Quercus sect. Protobalanus.

The Mendocin Indians considered the nuts of Quercus chrysolepis poisonous (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Protobalanus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. virginiana var. fusiformis Q. chrysolepis var. nana, Q. wilcoxii
Name authority Small: Bull. Torrey Bot. Club 28: 357. (1901) Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 173. (1854)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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