Quercus engelmannii |
Quercus wislizenii |
|
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Engelmann oak, Engelmann's oak |
interior live oak, Sierra live oak |
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Habit | Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed. | Trees or shrubs, evergreen, to 22 m. Bark nearly black, deeply furrowed with broad scaly ridges. |
Twigs | light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate. |
brown to red-brown, 1.5-3 mm diam., glabrous or sparsely pubescent. |
Buds | reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent; stipules persistent about terminal buds. |
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Leaves | blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded; surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous. |
blade circular to oblong, usually ovate, planar, 25-70 × 20-50 mm, base obtuse to cordate, margins entire or spinose with up to 16 awns, apex acute to rounded; surfaces abaxially and adaxially glabrous, veins little raised on either surface. |
Acorns | solitary or paired, subsessile or on peduncle to 5-6 mm; cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent; nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex. |
biennial; cup deeply and narrowly cup-shaped or U-shaped, 9-19 mm high × 7-18 mm wide, covering 1/3-1/2(-2/3) nut, outer surface glabrous to sparsely puberulent, inner surface glabrous or pubescent on innermost 1/3, occasionally uniformly pubescent, scales acute, tips loose; nut narrowly conic or ovoid to narrowly oblong, 21-44 × 8-14 mm, glabrous, scar diam. 2.5-7.5 mm. |
Cotyledons | connate. |
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Terminal | buds light chestnut brown to dark reddish brown, ovoid to conic, 3-9 mm, glabrous or with tuft of minute hairs at apex. |
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2n | = 24. |
= 24. |
Quercus engelmannii |
Quercus wislizenii |
|
Phenology | Flowering in spring. | Flowering late spring. |
Habitat | Oak woodlands, margins of chaparral, arroyos, slopes and bajadas | Valleys, slopes, and sand chaparral |
Elevation | 50-1200 m (200-3900 ft) | 300-1900 m (1000-6200 ft) |
Distribution |
CA; Mexico (Baja California)
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CA
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Discussion | Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment). On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Shrubs with oval leaves 25-38 mm and margins entire or deeply lobed-dentate may be treated as Quercus wislizenii var. frutescens. J. M. Tucker (1993) treated Q. parvula as a distinct species, distinguished from Q. wislizenii by its larger leaves (30-90 versus 20-50 mm), by the dull, olive-green, abaxial leaf surface (versus shiny, yellow-green), and by nuts that are abruptly tapered proximal to the middle (versus gradually tapered). Tucker recognized two varieties of Quercus parvula: Q. parvula var. parvula is a shrub of 1-3 m and Q. parvula var. shrevei is a tree less than 17 m. S. K. Langer (1993) recognized a third variety, Q. parvula var. tamalpaisensis,based on several small populations on or near Mount Tamalpais, differentiated primarily by having larger leaves (50-160 × 20-60 mm) with attenuate-dentate margins. Quercus wislizenii reportedly hybridizes with Q. agrifolia and Q. kelloggii (W. B. Brophy and D. R. Parnell 1974). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Quercus | Fagaceae > Quercus > sect. Lobatae |
Sibling taxa | ||
Synonyms | Q. parvula, Q. parvula var. shrevei, Q. parvula var. tamalpaisensis, Q. wislizenii var. frutescens | |
Name authority | Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889) | A. de Candolle: in A. P. de Candolle and A. L. P. de Candolle, Prodr. 16(2): 67. (1864) |
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