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Engelmann oak, Engelmann's oak

bastard oak, bastard white oak, Durand oak, Durand white oak

Habit Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed. Trees or shrubs, deciduous, to 15(-20) m, with solitary or multiple trunks.
Bark

gray to light brown, flaky to papery and exfoliating.

Twigs

light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate.

light gray or gray, 1-2(-3) mm diam., glabrous, rarely minutely puberulent.

Buds

reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent;

stipules persistent about terminal buds.

brown or reddish brown, broadly ovoid, 2-3 mm, essentially glabrous.

Leaves

blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded;

surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous.

blade oblong to oblanceolate, or narrowly rhomboid, or cuneiform, or rounded-3-dentate, (25-)30-120(-140) × (15-)25-60 mm, base acute, cuneate, attenuate-rounded, or obtuse, margins entire to irregularly toothed or moderately, sinuately lobed, flat, secondary veins ca. 7-11 on each side, apex broadly rounded, rarely attenuately narrowed or obscurely 3-lobed;

surfaces abaxially silvery or dull green, with scattered to crowded, minute, appressed-stellate, 8-10-rayed hairs, or glabrate or glabrous, especially in shade forms, adaxially green or dull green, glabrous.

Acorns

solitary or paired, subsessile or on peduncle to 5-6 mm;

cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent;

nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex.

solitary or paired, subsessile or on axillary peduncle to 1-7 mm;

cup saucer-shaped to shallowly cup-shaped, rarely deeper, 2-8 mm deep × 8-15(-20) mm wide, enclosing 1/8-1/4 nut, rarely more, base flat, rounded, or constricted, margin thin, scales closely appressed, grayish with reddish margins, ovate, flat, obtuse, not tuberculate;

nut light brown, depressed-ovoid to oblong, 7-15 × 7-12(-17) mm, glabrous.

Cotyledons

connate.

distinct.

2n

= 24.

Quercus engelmannii

Quercus sinuata

Phenology Flowering in spring.
Habitat Oak woodlands, margins of chaparral, arroyos, slopes and bajadas
Elevation 50-1200 m (200-3900 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment).

On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The question of the correct name for this species has persisted, with some authors rejecting the usage here in favor of Quercus durandii. Although no type material is extant, the original description of Q. sinuata is consistent with the concept presented here, as by W. W. Ashe (1916) and W. Trelease (1924), and inconsistent with any other oak from the broad area covered by Thomas Walter's Flora Caroliniana (1788).

The two varieties differ in habit, habitat, leaf size and lobing, and geographic range, and considerable variability exists within both varities as to the degree and density of silvery stellate-pubescence on the abaxial surface of the leaf. Sun leaves of both tend to have a higher proportion of silvery pubescence, and shade leaves and some individual trees tend to have more glabrate leaves, although evidence of flat-stellate trichomes is usually apparent. Plants with young, expanding leaves sometimes are mistaken for Quercus nigra, a member of the red oak group.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trees to 20 m, trunk usually solitary; moist bottomlands and riparian habitats.
var. sinuata
1. Shrubs or small trees to 3(–5) m, often forming thickets, trunks multiple from near ground; dry limestone hills.
var. breviloba
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Subordinate taxa
Q. sinuata var. breviloba, Q. sinuata var. sinuata
Synonyms Q. durandii
Name authority Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889) Walter: Fl. Carol., 235. (1788)
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