FNA: Quercus engelmannii vs. Quercus lobata

	Quercus engelmannii	Quercus lobata
	Engelmann oak, Engelmann's oak	California white oak, roble, valley oak
Habit	Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed.	Trees, deciduous, to 25(-35) m, usually with solitary trunks.
Bark		gray, scaly, deeply checkered in age.
Twigs	light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate.	yellowish, gray, occasionally reddish, 2-4 mm diam., densely or sparsely tomentulose.
Buds	reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent; stipules persistent about terminal buds.	yellowish or light brown, ovoid, (2-)3-5(-6) mm, apex occasionally acute, densely pubescent.
Leaves	blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded; surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous.	blade broadly obovate or elliptic, moderately to deeply lobed, (40-)50-100(-120) × 30-60(-75) mm, base rounded-attenuate, cuneate, or truncate, rarely subcordate, margins with sinuses usually reaching more than 1/2 distance to midrib, lobes oblong or spatulate, obtuse, rounded, or blunt, secondary veins 5-10 on each side, apex broadly rounded; surfaces abaxially whitish or light green, densely to sparsely covered with interlocking appressed or semi-erect, 8-10(-14)-rayed stellate hairs, adaxially dark green or grayish, glossy or somewhat scurfy because of sparse stellate hairs.
Acorns	solitary or paired, subsessile or on peduncle to 5-6 mm; cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent; nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex.	solitary or paired, subsessile; cup deeply cup-shaped, hemispheric or turbinate, rim thick, 10-30 mm deep × 14-30 mm wide, scales grayish or cream, more acute near rim, strongly and irregularly tuberculate, especially toward base of cup; nut light brown, oblong or fusiform, 30-60 × (12-)15-25 mm, tapering to acute or rounded apex.
Cotyledons	connate.	distinct.
2n	= 24.

	Quercus engelmannii	Quercus lobata
Phenology	Flowering in spring.	Flowering late winter–early spring.
Habitat	Oak woodlands, margins of chaparral, arroyos, slopes and bajadas	Valley floors and moderate slopes, open grasslands, savannah and oak woodlands, riparian areas in chaparral
Elevation	50-1200 m (200-3900 ft)	0-1700 m (0-5600 ft)
Distribution	CA; Mexico (Baja California) [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment). On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Mature trees of Quercus lobata are among the largest oaks of the United States. The species hybridizes with numerous other species, but the hybrids are not common in most parts of its range. On Santa Cruz and Santa Catalina islands, however, occur extensive and relatively stable populations that show intermediate characteristics with Q. pacifica (see treatment). The hybrids have been given the name Q. ×macdonaldii, and they differ from Q. lobata in the following: leaf sinuses reaching less than half the distance to the midrib; leaves usually smaller, the lobes often more acute and brighter green; and acorns smaller, with more acute apices. The Yuri used Quercus lobata in the treatment of diarrhea (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms		Q. hindsii, Q. lobata var. hindsii, Q. longiglanda
Name authority	Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889)	Nee.: Anales Ci. Nat. 3: 277. (1801)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus lobata

Engelmann oak, Engelmann's oak

California white oak, roble, valley oak

Habit

Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed.

Trees, deciduous, to 25(-35) m, usually with solitary trunks.

Bark

gray, scaly, deeply checkered in age.

Twigs

light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate.

yellowish, gray, occasionally reddish, 2-4 mm diam., densely or sparsely tomentulose.

Buds

reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent;

stipules persistent about terminal buds.

yellowish or light brown, ovoid, (2-)3-5(-6) mm, apex occasionally acute, densely pubescent.

Leaves

blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded;

surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous.

blade broadly obovate or elliptic, moderately to deeply lobed, (40-)50-100(-120) × 30-60(-75) mm, base rounded-attenuate, cuneate, or truncate, rarely subcordate, margins with sinuses usually reaching more than 1/2 distance to midrib, lobes oblong or spatulate, obtuse, rounded, or blunt, secondary veins 5-10 on each side, apex broadly rounded;

surfaces abaxially whitish or light green, densely to sparsely covered with interlocking appressed or semi-erect, 8-10(-14)-rayed stellate hairs, adaxially dark green or grayish, glossy or somewhat scurfy because of sparse stellate hairs.

Acorns

solitary or paired, subsessile or on peduncle to 5-6 mm;

cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent;

nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex.

solitary or paired, subsessile;

cup deeply cup-shaped, hemispheric or turbinate, rim thick, 10-30 mm deep × 14-30 mm wide, scales grayish or cream, more acute near rim, strongly and irregularly tuberculate, especially toward base of cup;

nut light brown, oblong or fusiform, 30-60 × (12-)15-25 mm, tapering to acute or rounded apex.

Cotyledons

connate.

distinct.

= 24.

Quercus engelmannii

Quercus lobata

Phenology

Flowering in spring.

Flowering late winter–early spring.

Habitat

Oak woodlands, margins of chaparral, arroyos, slopes and bajadas

Valley floors and moderate slopes, open grasslands, savannah and oak woodlands, riparian areas in chaparral

Elevation

50-1200 m (200-3900 ft)

0-1700 m (0-5600 ft)

Distribution

CA; Mexico (Baja California)

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment).

On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mature trees of Quercus lobata are among the largest oaks of the United States. The species hybridizes with numerous other species, but the hybrids are not common in most parts of its range. On Santa Cruz and Santa Catalina islands, however, occur extensive and relatively stable populations that show intermediate characteristics with Q. pacifica (see treatment). The hybrids have been given the name Q. ×macdonaldii, and they differ from Q. lobata in the following: leaf sinuses reaching less than half the distance to the midrib; leaves usually smaller, the lobes often more acute and brighter green; and acorns smaller, with more acute apices.

The Yuri used Quercus lobata in the treatment of diarrhea (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. hindsii, Q. lobata var. hindsii, Q. longiglanda

Name authority

Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889)

Nee.: Anales Ci. Nat. 3: 277. (1801)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus engelmannii

Quercus lobata

Quercus engelmannii

Quercus lobata