Quercus engelmannii |
Quercus georgiana |
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Engelmann oak, Engelmann's oak |
Georgia oak, stone mountain oak |
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Habit | Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed. | Trees, deciduous, to 15 m. Bark gray to light brown, scaly. |
Twigs | light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate. |
deep red, 1-2 mm diam., glabrous. |
Buds | reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent; stipules persistent about terminal buds. |
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Leaves | blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded; surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous. |
blade broadly ovate to elliptic or obovate, 40-130 × 20-90 mm, base cuneate to obtuse, margins with 3-5(-7) oblong lobes and up to 10 awns, apex acute; surfaces abaxially glabrous except for conspicuous axillary tufts of tomentum, veins raised, adaxially planar, glabrous. |
Acorns | solitary or paired, subsessile or on peduncle to 5-6 mm; cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent; nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex. |
biennial; cup thin, saucer-shaped, 4-6 mm high × 9-14 mm wide, covering 1/3 nut, outer surface puberulent, inner surface glabrous or with a few hairs around scar, scale tips appressed, acute; nut globose or ovoid, 9-14 × 9-14 mm, glabrous, scar diam. 4-7.5 mm. |
Cotyledons | connate. |
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Terminal | buds red-brown, ovoid to subconic, 2.5-5 mm, glabrous or scales somewhat ciliate. |
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2n | = 24. |
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Quercus engelmannii |
Quercus georgiana |
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Phenology | Flowering in spring. | Flowering spring. |
Habitat | Oak woodlands, margins of chaparral, arroyos, slopes and bajadas | Granitic outcrops and dry slopes and knolls |
Elevation | 50-1200 m (200-3900 ft) | 50-500 m (200-1600 ft) |
Distribution |
CA; Mexico (Baja California)
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AL; GA; SC
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Discussion | Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment). On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Quercus georgiana reportedly hybridizes with Q. marilandica (= Q. ×smallii Trelease) and Q. nigra, although D. M. Hunt (1989) has questioned the validity of the former report. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Quercus | Fagaceae > Quercus > sect. Lobatae |
Sibling taxa | ||
Name authority | Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889) | M. A. Curtis: Amer. J. Sci. Arts, ser. 2, 7: 406. (1849) |
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