FNA: Quercus engelmannii vs. Quercus douglasii

	Quercus engelmannii	Quercus douglasii
	Engelmann oak, Engelmann's oak	blue oak
Habit	Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed.	Trees, deciduous, with single trunks, sometimes with few to several trunks.
Bark		gray, scaly.
Twigs	light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate.	reddish or yellowish, ca. 2 mm, densely or sparsely puberulent, occasionally glabrate with age.
Buds	reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent; stipules persistent about terminal buds.	reddish brown, rarely yellowish, broadly ovoid to rarely subglobose, (2-)3-5 mm; scales glabrous except for ciliate margins, sometimes sparsely or densely pubescent.
Leaves	blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded; surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous.	blade obovate or elliptic, oblong or oblanceolate, (20-)40-60(-80) × (15-)20-30(-40) mm, base rounded-attenuate or rounded, rarely cuneate, margins shallowly lobed or irregularly toothed, sometimes entire, lobes mucronate or rounded, secondary veins 6-10 on each side, apex rounded, rarely moderately acute; surfaces abaxially light green or blue-green, waxy, with scattered to crowded, semi-erect, (2-)4-6(-8)-rayed stellate hairs usually 0.2-0.6 mm diam. or larger, adaxially blue-green, glaucous or grayish, vestiture similar to abaxial surface.
Acorns	solitary or paired, subsessile or on peduncle to 5-6 mm; cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent; nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex.	subsessile, solitary; cup hemispheric or cup-shaped, rarely deeper, 5-10 mm deep × 10-15 mm wide, enclosing only base of nut, scales thin and not tuberculate to strongly and irregularly tuberculate, particularly toward base of cup; nut thin-walled, fusiform or subcylindric, 20-30 × 10-16 mm.
Cotyledons	connate.	distinct.
2n	= 24.	= 24.

	Quercus engelmannii	Quercus douglasii
Phenology	Flowering in spring.	Flowering late winter–spring.
Habitat	Oak woodlands, margins of chaparral, arroyos, slopes and bajadas	Oak woodlands, margins of chaparral and grasslands
Elevation	50-1200 m (200-3900 ft)	0-1200 m (0-3900 ft)
Distribution	CA; Mexico (Baja California) [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment). On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Populations of Quercus douglasii in coastal southern California and on the Channel Islands consist of small stands or solitary individuals often associated with Q. lobata and scrub oaks. Some doubt exists as to whether some or all of those populations are natural stands or are historical introductions near Native American settlements. Along the canyons of Santa Barbara County, putative hybrids between Q. douglasii and Q. dumosa are referred to Q. ×kinselae (C. H. Muller) Nixon. In the interior Coast Ranges of California are found numerous populations that are intermediate in form between Q. douglasii and Q. john-tuckeri (= Q. turbinella var. californica). This appears to be an area of secondary contact, and the two species remain distinct in nearby populations. Because of the widespread nature of the intermediates, following Tucker's extensive studies they can be conveniently referred to as the nothospecies Q. ×alvordiana Eastwood. The plants tend to be shrubs to small trees, with somewhat more spinose leaves than Q. douglasii and fruit similar to those of the latter species. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms		Q. douglasii var. ransomii, Q. ransomii
Name authority	Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889)	Hooker & Arnott: Bot. Beechey Voy. 9: 391. (1840)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus douglasii

Engelmann oak, Engelmann's oak

blue oak

Habit

Trees, subevergreen, to 10 m. Bark gray or whitish, closely furrowed.

Trees, deciduous, with single trunks, sometimes with few to several trunks.

Bark

gray, scaly.

Twigs

light brown, 1-1.5 mm diam., densely or sparsely stellate-tomentose, soon glabrate.

reddish or yellowish, ca. 2 mm, densely or sparsely puberulent, occasionally glabrate with age.

Buds

reddish brown, subspheric to broadly ovoid, 1-2 mm, glabrous or basal scales pubescent;

stipules persistent about terminal buds.

reddish brown, rarely yellowish, broadly ovoid to rarely subglobose, (2-)3-5 mm;

scales glabrous except for ciliate margins, sometimes sparsely or densely pubescent.

Leaves

blade oblong to elliptic, occasionally lanceolate or ovate, (20-)30-60(-80) × (5-)10-20(-25) mm, base cuneate to cordate, margins entire, undulate, sometimes irregularly toothed, especially toward apex, secondary veins 7-8(-10) on each side, branched, apex acute or broadly rounded;

surfaces abaxially blue-green or pale green, densely and loosely glandular-tomentose, quickly glabrate or persistently floccose, especially about base of midrib, at maturity strongly glaucous, adaxially gray-green or pale green, bluish green or glaucous.

blade obovate or elliptic, oblong or oblanceolate, (20-)40-60(-80) × (15-)20-30(-40) mm, base rounded-attenuate or rounded, rarely cuneate, margins shallowly lobed or irregularly toothed, sometimes entire, lobes mucronate or rounded, secondary veins 6-10 on each side, apex rounded, rarely moderately acute;

surfaces abaxially light green or blue-green, waxy, with scattered to crowded, semi-erect, (2-)4-6(-8)-rayed stellate hairs usually 0.2-0.6 mm diam. or larger, adaxially blue-green, glaucous or grayish, vestiture similar to abaxial surface.

Acorns

solitary or paired, subsessile or on peduncle to 5-6 mm;

cup cup-shaped or shallowly cup-shaped, 8-10 mm deep × 10-15 mm wide, enclosing 1/3 nut, scales 1.5-3 mm wide, strongly and regularly tuberculate near base of cup, gray-pubescent;

nut light brown, ovoid or oblong, 15-25 × 12-14 mm, glabrate or puberulent about apex.

subsessile, solitary;

cup hemispheric or cup-shaped, rarely deeper, 5-10 mm deep × 10-15 mm wide, enclosing only base of nut, scales thin and not tuberculate to strongly and irregularly tuberculate, particularly toward base of cup;

nut thin-walled, fusiform or subcylindric, 20-30 × 10-16 mm.

Cotyledons

connate.

distinct.

= 24.

Quercus engelmannii

Quercus douglasii

Phenology

Flowering in spring.

Flowering late winter–spring.

Habitat

Oak woodlands, margins of chaparral, arroyos, slopes and bajadas

Oak woodlands, margins of chaparral and grasslands

Elevation

50-1200 m (200-3900 ft)

0-1200 m (0-3900 ft)

Distribution

CA; Mexico (Baja California)

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and generally more tuberculate than those of Q. oblongifolia, and the scales are usually larger. Based on available samples, the nuts of Q. engelmannii are consistently larger than those of Q. oblongifolia, apparently with little, if any, overlap in diameter. Considerably more variation occurs within Q. engelmannii in leaf form, possibly reflecting introgression from other white oak species such as Q. cornelius-mulleri, Q. dumosa, and Q. durata (see treatment).

On Catalina Island, Quercus engelmannii is known only from a small grove of trees. Putative hybrids between Q. engelmannii and Q. cornelius-mulleri are common in areas of contact between the two species in Riverside and San Diego counties in southern California. Such a population was the basis for Q. acutidens Torrey [Q. dumosa var. acutidens (Torrey) Wenzig]. Other names applied to those populations are Q. macdonaldii var. elegantula Greene and Q. dumosa var. elegantula (Greene) Jepson. Variable in leaf form and stature, those intermediates form extensive populations and are probably best disposed of under the name Q. ×acutidens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Quercus douglasii in coastal southern California and on the Channel Islands consist of small stands or solitary individuals often associated with Q. lobata and scrub oaks. Some doubt exists as to whether some or all of those populations are natural stands or are historical introductions near Native American settlements. Along the canyons of Santa Barbara County, putative hybrids between Q. douglasii and Q. dumosa are referred to Q. ×kinselae (C. H. Muller) Nixon. In the interior Coast Ranges of California are found numerous populations that are intermediate in form between Q. douglasii and Q. john-tuckeri (= Q. turbinella var. californica). This appears to be an area of secondary contact, and the two species remain distinct in nearby populations. Because of the widespread nature of the intermediates, following Tucker's extensive studies they can be conveniently referred to as the nothospecies Q. ×alvordiana Eastwood. The plants tend to be shrubs to small trees, with somewhat more spinose leaves than Q. douglasii and fruit similar to those of the latter species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. douglasii var. ransomii, Q. ransomii

Name authority

Greene: Ill. W. Amer. Oaks 1: 33, plate 17. (1889)

Hooker & Arnott: Bot. Beechey Voy. 9: 391. (1840)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Quercus engelmannii

Quercus douglasii

Quercus engelmannii

Quercus douglasii