FNA: Quercus ellipsoidalis vs. Quercus arizonica

	Quercus ellipsoidalis	Quercus arizonica
	Hill's oak, Jack oak, northern pin oak	Arizona oak, Arizona white oak
Habit	Trees, deciduous, to 20 m; lower trunk often with stubs of dead branches.	Trees, evergreen or subevergreen, small to moderate-sized trees, rarely to 18 m. Bark scaly.
Bark	dark gray-brown, shallowly fissured, inner bark orangish.
Twigs	dark reddish brown, (1-)1.5-3 mm diam., glabrous.	yellowish, 1.5-2.5 mm diam., persistently felty-tomentose, eventually dingy gray.
Buds		dull russet-brown, ovoid, distally subacute or rounded, 3 mm, sparsely pubescent or glabrate.
Leaves	blade elliptic, 70-130 × 50-100 mm, base obtuse to truncate, margins with 5-7 deep lobes and 15-55 awns, lobes distally expanded, sinuses usually extending more than 1/2 distance to midrib, apex acute; surfaces abaxially glabrous except for minute axillary tufts of tomentum, adaxially glossy light green, glabrous, secondary veins raised on both surfaces.	blade elliptic or oblong to narrowly obovate or oblanceolate, planar or moderately convex, to (30-)40-80(-90) × 15-30 mm, thick and leathery, usually stiff, base cordate or rounded and weakly cordate, margins entire or coarsely toothed especially near apex, cartilaginously revolute, teeth mucronate-tipped, obscure or prominent, secondary veins ca. 7-11 on each side, branching, passing into teeth when present, apex acute to usually obtuse or broadly rounded; surfaces abaxially dull, sparsely pubescent or subtomentose with curly branched hairs, reticulate from prominent, raised secondary veins, usually glaucous where exposed, adaxially dark or bluish green, moderately lustrous, sparsely and minutely stellate-pubescent, secondary veins slightly raised or prominent within depressions or impressed.
Acorns	biennial; cup narrowly turbinate to deeply cup-shaped, 6-11 mm high × 10-19 mm wide, covering 1/3-1/2 nut, outer surface reddish brown, puberulent, inner surface light brown, glabrous, rarely with ring of pubescence around scar, scales with straight or slightly concave margins, tips tightly appressed, obtuse or acute; nut ellipsoid to ovoid, rarely subglobose, 10-20 × 9-15 mm, occasionally striate, glabrous, occasionally with 1 or more faint rings of fine pits at apex, scar diam. 4-8 mm.	solitary or paired, subsessile, occasionally on peduncle to 15 mm; cup hemispheric or cup-shaped, 5-10(-15) mm deep × 10-15 mm wide, enclosing ca. 1/2 nut, base rounded, margin rather coarse, scales cream to brown, broadly ovate, evenly and strongly tuberculate, tomentose, tips closely appressed; nut light brown, ovoid or oblong, 8-12 mm, nearly glabrous.
Cotyledons		connate.
Terminal	buds dark reddish brown, ovoid, 3-5 mm, often conspicuously 5-angled in cross section, usually silvery- or tawny-pubescent toward apex.

	Quercus ellipsoidalis	Quercus arizonica
Phenology	Flowering spring.	Flowering spring.
Habitat	Dry sandy sites, rarely on moderately mesic slopes or uplands	Oak and pinyon woodlands, margins of chaparral, arroyos
Elevation	150-500 m (500-1600 ft)	1300-2500(-3000) m (4300-8200(-9800) ft)
Distribution	IA; IL; IN; MI; MN; OH; WI; ON [WildflowerSearch map] [BONAP county map]	AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, and Sonora) [WildflowerSearch map] [BONAP county map]
Discussion	In many treatments (e.g., E. G. Voss 1972+, vol. 2), Quercus ellipsoidalis is included in Q. coccinea. Variation in fruit morphology has led to recognition of several formae (W. Trelease 1919; see also R. J. Jensen 1986) and one variety (Q. ellipsoidalis var. kaposianensis, based on specimens from St. Paul, Minnesota, in which the cup tightly encloses the nut for two-thirds its length at maturity). Quercus ellipsoidalis reportedly hybridizes with Q. rubra and Q. velutina. The Menominee used Quercus ellipsoidalis medicinally to treat suppressed menses caused by cold (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Some of the specimens previously referred to Quercus endemica by C. H. Muller belong here instead. Putative hybrids between Quercus arizonica and Q. grisea (= Q. ×organensis Trelease) are problematic in local areas of contact from southeastern Arizona to western Texas. These intermediates tend to have narrower leaves than Q. arizonica, with moderately reticulate patterns of venation, and more densely hairy leaves. Quercus arizonica and Q. grisea are amply distinct elsewhere, including large areas in northern Mexico, and they appear to be more closely related to other species than to one another (e.g., Q. arizonica with Q. oblongifolia and Q. laeta Liebmann, and Q. grisea with Q. mohriana and Q. microphylla Née). Thus, Q. arizonica and Q. grisea are best treated as distinct species that hybridize, and not as conspecific populations. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Lobatae	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. ellipsoidalis var. kaposianensis	Q. sacame
Name authority	E. J. Hill: Bot. Gaz. 27: 204, plates 2, 3. (1899)	Sargent: Gard. & Forest 8: 92. (1895)
Web links	Local Web sites: IL Wildflowers, MI Flora, MN Wildflowers WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus arizonica

Hill's oak, Jack oak, northern pin oak

Arizona oak, Arizona white oak

Habit

Trees, deciduous, to 20 m; lower trunk often with stubs of dead branches.

Trees, evergreen or subevergreen, small to moderate-sized trees, rarely to 18 m. Bark scaly.

Bark

dark gray-brown, shallowly fissured, inner bark orangish.

Twigs

dark reddish brown, (1-)1.5-3 mm diam., glabrous.

yellowish, 1.5-2.5 mm diam., persistently felty-tomentose, eventually dingy gray.

Buds

dull russet-brown, ovoid, distally subacute or rounded, 3 mm, sparsely pubescent or glabrate.

Leaves

blade elliptic, 70-130 × 50-100 mm, base obtuse to truncate, margins with 5-7 deep lobes and 15-55 awns, lobes distally expanded, sinuses usually extending more than 1/2 distance to midrib, apex acute;

surfaces abaxially glabrous except for minute axillary tufts of tomentum, adaxially glossy light green, glabrous, secondary veins raised on both surfaces.

blade elliptic or oblong to narrowly obovate or oblanceolate, planar or moderately convex, to (30-)40-80(-90) × 15-30 mm, thick and leathery, usually stiff, base cordate or rounded and weakly cordate, margins entire or coarsely toothed especially near apex, cartilaginously revolute, teeth mucronate-tipped, obscure or prominent, secondary veins ca. 7-11 on each side, branching, passing into teeth when present, apex acute to usually obtuse or broadly rounded;

surfaces abaxially dull, sparsely pubescent or subtomentose with curly branched hairs, reticulate from prominent, raised secondary veins, usually glaucous where exposed, adaxially dark or bluish green, moderately lustrous, sparsely and minutely stellate-pubescent, secondary veins slightly raised or prominent within depressions or impressed.

Acorns

biennial;

cup narrowly turbinate to deeply cup-shaped, 6-11 mm high × 10-19 mm wide, covering 1/3-1/2 nut, outer surface reddish brown, puberulent, inner surface light brown, glabrous, rarely with ring of pubescence around scar, scales with straight or slightly concave margins, tips tightly appressed, obtuse or acute;

nut ellipsoid to ovoid, rarely subglobose, 10-20 × 9-15 mm, occasionally striate, glabrous, occasionally with 1 or more faint rings of fine pits at apex, scar diam. 4-8 mm.

solitary or paired, subsessile, occasionally on peduncle to 15 mm;

cup hemispheric or cup-shaped, 5-10(-15) mm deep × 10-15 mm wide, enclosing ca. 1/2 nut, base rounded, margin rather coarse, scales cream to brown, broadly ovate, evenly and strongly tuberculate, tomentose, tips closely appressed;

nut light brown, ovoid or oblong, 8-12 mm, nearly glabrous.

Cotyledons

connate.

Terminal

buds dark reddish brown, ovoid, 3-5 mm, often conspicuously 5-angled in cross section, usually silvery- or tawny-pubescent toward apex.

Quercus ellipsoidalis

Quercus arizonica

Phenology

Flowering spring.

Habitat

Dry sandy sites, rarely on moderately mesic slopes or uplands

Oak and pinyon woodlands, margins of chaparral, arroyos

Elevation

150-500 m (500-1600 ft)

1300-2500(-3000) m (4300-8200(-9800) ft)

Distribution

IA; IL; IN; MI; MN; OH; WI; ON

[WildflowerSearch map]

[BONAP county map]

AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, and Sonora)

[WildflowerSearch map]

[BONAP county map]

Discussion

In many treatments (e.g., E. G. Voss 1972+, vol. 2), Quercus ellipsoidalis is included in Q. coccinea. Variation in fruit morphology has led to recognition of several formae (W. Trelease 1919; see also R. J. Jensen 1986) and one variety (Q. ellipsoidalis var. kaposianensis, based on specimens from St. Paul, Minnesota, in which the cup tightly encloses the nut for two-thirds its length at maturity).

Quercus ellipsoidalis reportedly hybridizes with Q. rubra and Q. velutina.

The Menominee used Quercus ellipsoidalis medicinally to treat suppressed menses caused by cold (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some of the specimens previously referred to Quercus endemica by C. H. Muller belong here instead.

Putative hybrids between Quercus arizonica and Q. grisea (= Q. ×organensis Trelease) are problematic in local areas of contact from southeastern Arizona to western Texas. These intermediates tend to have narrower leaves than Q. arizonica, with moderately reticulate patterns of venation, and more densely hairy leaves. Quercus arizonica and Q. grisea are amply distinct elsewhere, including large areas in northern Mexico, and they appear to be more closely related to other species than to one another (e.g., Q. arizonica with Q. oblongifolia and Q. laeta Liebmann, and Q. grisea with Q. mohriana and Q. microphylla Née). Thus, Q. arizonica and Q. grisea are best treated as distinct species that hybridize, and not as conspecific populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Lobatae

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. ellipsoidalis var. kaposianensis

Q. sacame

Name authority

E. J. Hill: Bot. Gaz. 27: 204, plates 2, 3. (1899)

Sargent: Gard. & Forest 8: 92. (1895)

Web links

Local Web sites: IL Wildflowers, MI Flora, MN Wildflowers
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Quercus ellipsoidalis

Quercus arizonica

Quercus ellipsoidalis

Quercus arizonica