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coastal sage scrub oak, Nuttall's scrub oak, scrub oak

escarpment live oak, live oak, plateau oak, Texas live oak

Habit Shrubs, subevergreen, 1-2(-2.5) m, dense, divaricately branching, leaves brittle, often falling when branches disturbed. Trees, sometimes shrubs, subevergreen, trees to 25 m, shrubs often forming large clonal stands.
Bark

smooth when young, eventually scaly.

dark brown or black, scaly.

Twigs

reddish or grayish, 1-1.5(-2) mm diam., glabrous or sparsely stellate-pubescent, soon glabrate.

light gray, 1.5-3 mm diam., tomentulose, tomentulum often persistent in age.

Buds

reddish brown, globose or ovoid, 1-2 mm, glabrous, rarely puberulent near apex.

reddish or dark brown, subglobose or ovate, 1.5-3 mm;

scale margins glabrous or puberulent.

Leaves

blade undulate or strongly to moderately cupped, occasionally subplanar, 10-20(-25) × 6-15(-20) mm, base cordate or angular-cordate, margins irregularly spinose-toothed or shallowly lobed, rarely entire, often somewhat revolute, secondary veins 3-5(-6) on each side, irregularly branched, apex rounded or spinose-acute;

surfaces abaxially sparsely to densely covered with erect, curly, (2-)4(-6)-rayed fasciculate hairs to 0.5 mm, felty to touch in young leaves, adaxially glossy green, glabrate or with scattered stellate hairs, secondary veins somewhat impressed, puberulent.

blade oblong-elliptic to narrowly ovate or lanceolate, sometimes obovate, ± planar, (10-)35-90(-150) × (15-)20-40(-85) mm, base rounded to truncate or cordate, rarely cuneate, margins minutely revolute or flat, entire or irregularly 1-3 toothed on each side, teeth mucronate (rarely spinose in suckers or juveniles), secondary veins obscure, 8-10 on each side, apex obtuse-rounded or acute;

surfaces abaxially whitish or glaucous, densely covered with minute, appressed, fused-stellate hairs, light green and glabrate in shade leaves, adaxially dark or light green, glossy, glabrous or with minute, scattered, stellate hairs.

Acorns

solitary or paired, subsessile;

cup reddish, deeply cup-shaped, 5-8 mm deep × 8-15 mm wide, enclosing 1/3 nut or less, scales long-acute, moderately or scarcely tuberculate at base;

nut fusiform or subcylindric, 10-20(-30) × 5-10(-12) mm, apex acute.

1-3, on peduncle 3-30 mm;

cup funnel-shaped, hemispheric, or deeply goblet-shaped, 8-15 mm deep × 6-12(-15) mm wide, base often constricted, scales whitish or grayish, thickened basally, keeled, acute-attenuate, tomentulose, tips reddish, glabrous or puberulent;

nut dark brown, often with light brown longitudinal stripes, subfusiform and acute to narrowly barrel-shaped, rarely distally rounded, (17-)20-30(-33) × 8-15 mm, glabrous.

Cotyledons

distinct.

connate.

Quercus dumosa

Quercus fusiformis

Phenology Flowering spring. Flowering spring.
Habitat Open chaparral, coastal sage scrub Hills, grasslands, scrublands, open woodlands, oak-juniper woodland, and margins of thorn scrub, often on limestone or deep calcareous loams, sometimes on granular sand or gravel
Elevation 0-300 m (0-1000 ft) 0-1200 m (0-3900 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX; Mexico (Coahuila, Nuevo León, and Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

The name Quercus dumosa has been applied to virtually all scrub oaks in the white oak group of central and southern California and adjacent Baja California. Through the years, and following independent studies by various authors, the concept of this species has gradually narrowed from the original, which included plants here segregated as Q. turbinella, Q. john-tuckeri, Q. cornelius-mulleri, Q. berberidifolia, and Q. pacifica. In degree and constancy, the differences among these species are similar to those separating other commonly recognized tree species of the western United States. The majority of populations referred to Q. dumosa in recent treatments are now included in Q. berberidifolia (see treatment). All of the scrub oaks have a striking superficial similarity because of their shrubby habit and small, often spiny leaves; they differ dramatically in leaf and twig vestiture and acorn form. The concept of Q. dumosa presented here limits it to populations of scraggly shrubs with short petioles, cordate leaf bases, erect curly trichomes on the abaxial leaf surface, and narrow acute acorns that occur at low elevations almost always within sight of the ocean. Because these locations are typically prime real estate, the species, which probably never was common, is highly at risk. It rarely comes into contact with other white oaks because of its low elevation and dry habitat preference; putative hybrids are known, however, with Q. engelmannii and Q. lobata. Some populations of Q. berberidifolia from higher elevations near populations of Q. dumosa show signs of introgression.

Named hybrids include Quercus ×kinselae (C. H. Muller) Nixon & C. H. Muller (= Q. dumosa Nuttall × Q. lobata Née) and Q. dumosa Nuttall var. kinselae C. H. Muller (= Q. dumosa × Q. engelmannii).

The Luisenos used gall nuts from Quercus dumosa medicinally for sores and wounds and as an astringent (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The difficulty in distinguishing Texas populations of Quercus fusiformis from Q. virginiana is reflected in a variety of taxonomic treatments, including reducing Q. fusiformis to varietal rank under Q. virginiana. The latter disposition is problematic, however, because Q. fusiformis in northeastern Mexico is amply distinct from Q. virginiana and appears to be more closely related to Q. brandegei Goldmann, an endemic of Baja California, Mexico. Thus, here we assume that the intergradation of Q. virginiana and Q. fusiformis is a result of secondary contact, and is not primary clinal variation. Under this interpretation, Q. virginiana in typical form extends into Texas only as far west as the Brazos River drainage along the coast from there to the escarpment of the Edwards Plateau; most populations elsewhere are either intermediate between the two species or show greater affinity with Q. fusiformis. On the Edwards Plateau, the live oak populations are small trees forming rhizomatous copses (shinneries) and having mostly acute acorns.

Populations of live oak on deep sands in south Texas differ from typical Quercus fusiformis in having broader, more rounded leaves, often with the secondary veins somewhat impressed abaxially, and relatively blunt, barrel-shaped acorns. These characteristics suggest introgresion from the Mexican-Central American species Q. oleoides Schlechtendal & Chamisso, which in its typical form reaches north only as far as southern Tamaulipas, Mexico. The name Q. oleoides var. quaterna C. H. Muller has been applied to what is apparently a shrub form of one of these Q. fusiformis × Q. oleoides hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Quercus Fagaceae > Quercus > sect. Quercus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. virginiana var. fusiformis
Name authority Nuttall: N. Amer. Sylv. 1: 7. (1842) Small: Bull. Torrey Bot. Club 28: 357. (1901)
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